Above all, Gärtner put an end to the blunder of regarding dry
indehiscent fruits as naked seeds, by rightly defining the pericarp
as in all cases the ripened wall of the ovary, and by considering its
strong or weak construction, its dry or pulpy condition, as a secondary
matter. It is obvious that the whole theory of the flower was thus
placed upon a better basis, since dry indehiscent fruits may come from
inferior or superior ovaries. But Gärtner’s theory of the seed is
one of his most valuable contributions to the science. After careful
consideration of the seed-envelopes, he submitted the inner portion
(nucleus) enclosed by them to a searching comparative examination;
he correctly distinguished the endosperm from the cotyledons, and
described the variations in its form and position. This was the more
needful, since Linnaeus had denied the existence of an ‘albumen’ in
plants, which Grew had already recognised and so named; to Linnaeus
it appeared to be of no use to the seed. Though Gärtner speaks of the
cotyledons as uniting with the embryo to form the nucleus of the seed,
yet his account shows that he regarded them as outgrowths of the embryo
itself. The uncertainty which still existed in the interpretation of
the parts of the seed is shown even in Gärtner by his curious notion of
a ‘vitellus,’ which in fact takes in everything that he was unable to
explain aright inside the seed; for instance, he makes the scutellum
in grasses, and even the cotyledonary bodies of Zamia a vitellus, and
applies the same name to the whole contents of the spores of Seaweeds,
Mosses, and Ferns. In spite of the striking defects connected with
this mistaken notion in his theory of the seed, his views far surpass
in clearness and consistency all that had hitherto been taught on the
subject. His giving the term embryo to that part of the seed which is
capable of development was also an advance in respect of logic and
morphology, in spite of his mistake in not admitting the cotyledons
which are attached to the embryo into the conception; this, however,
could easily be corrected at a later time. What Gärtner now named
the embryo, had been up to his time called the ‘corculum seminis,’
especially by Linnaeus and Jussieu; it was evidently thought that
Cesalpino’s phraseology was thus retained; but he, as we have seen,
understood by the words ‘cor seminis’ the spot where the cotyledons
spring from the germ, which spot he wrongly took for the meeting-point
of root and stem and the seat of the soul of the plant. And so at last
after two hundred years the word disappeared from use, which might
have reminded the botanist of Cesalpino’s views respecting the soul of
plants.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account