De Candolle published his morphology or doctrine of symmetry and his
theory of classification together in a book which appeared first
in 1813, with the title, ‘Théorie Élémentaire de la botanique ou
exposition des principes de la classification naturelle et de l’art de
décrire et d’étudier les végétaux,’ and again in 1819 in an improved
and enlarged edition. The second edition will be the one referred to in
the further account of his views. The second chapter of the second book
concerns us most at present. After alluding to the fact, that anatomy
and physiology are concerned with the structure of the individual
organ only so far as the power to fulfil its proper function depends
on the structure, he points out that the physiological point of view
is no longer sufficient when we are engaged in comparing the organs of
different plants. Though it is true that the function of the organs
is the most important for the life and permanence of the individual,
yet we find these functions modified in the case of homologous organs
in different plants; for the natural classification we must take into
consideration only the entire system of organisation, that is, the
symmetry of the organs. All organisms of a kingdom, he continues, have
the same functions with slight modifications; the immense amount of
variation in systematically different species depends therefore only
on the way in which the general symmetry of structure varies. This
symmetry of the parts, the discovery of which is the great object
in the investigation of nature, is nothing more than the sum total
(l’ensemble) of the positional relations of the parts. Whenever these
relations (disposition) are regulated according to the same plan,
the organisms exhibit a certain general resemblance to one another,
independently of the form of the organs in detail; when this general
resemblance is perceived, without any attempt to give any account of
it in the detail, we have what has been called habitual relationship;
but it is the task of the doctrine of symmetry to resolve this likeness
of habit into its elements, and to explain its causes. Without this
study of symmetry it may easily happen that two different kinds of
symmetry may be supposed to be alike, because they seem outwardly
alike to our senses, just as forms of crystals of different systems
may be confounded together for want of careful examination; the chief
thing is to know the plan of symmetry in every class of plants,
and the study of this is the foundation of every theory of natural
affinities. But success in this study depends on the certainty with
which organs are distinguished, and the distinguishing them must be
independent of changes of form, size, and function. He then shows that
the difficulties in the morphological comparison of organs, or, as we
should now say, in the establishing the homology, are due to three
causes; abortion, degeneration, and adherence (adhérence). These three
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