and Endlicher in Germany, and that of Brongniart in France. After
various earlier attempts he proposed a system in 1845, in which, as
in Brongniart’s arrangement, the Cryptogams are characterised as
asexual or flowerless plants, the Phanerogams as sexual or flowering
plants; the former are divided into Thallogens and Acrogens, the
Phanerogams into five classes; (1) Rhizogens (Rafflesiaceae, Cytineae,
Balanophorae); (2) Endogens (parallel-nerved Monocotyledons); (3)
Dictyogens (net-veined Monocotyledons); (4) Gymnogens (Gymnosperms);
(5) Exogens (Dicotyledons). This classification is one of the most
unfortunate that were ever attempted; the systematic value of the
Rhizogens is much overrated on account of their striking habit; the
Monocotyledons are separated into two classes on the strength of an
unimportant mark. The characters assigned to all these groups are on
the whole thoroughly faulty.
These systems have been selected for notice from among many others,
because they attained an extended notoriety and importance from the
circumstance that their authors, Brongniart excepted, made them the
occasions of comprehensive descriptions of the whole vegetable kingdom,
and again because it would be superfluous for our present purpose to
bestow a closer consideration on the systems of less eminent men.
Whoever desires further information on the matter will find it in the
introduction to Lindley’s ‘Vegetable Kingdom’ of 1853.
If we consider the principles and points of view adopted in these
systems, one thing especially strikes us, that, except in the case
of Bartling, physiologico-anatomical marks were employed along with
morphological ones to characterise the primary divisions; their authors
fell into the mistake committed by De Candolle, and unfortunately
these very marks rested in part or wholly on misapprehensions, as in
Endlicher’s division into Acrobrya, etc., and Lindley’s classes of
Rhizogens and Dictyogens. It was still more unfortunate that individual
systematists obstinately refused to accept well authenticated facts,
which it is true had not been discovered by systematists, but were
nevertheless of the highest value for the system. It is scarcely
credible that Lindley in 1845, and again in 1853, maintained the
distinction between endogenous and exogenous growth in stems, though
Hugo von Mohl had in 1831 produced decisive proof that this distinction
laid down by Desfontaines and adopted by De Candolle had no real
existence. The same was the case with the characters of the Cryptogams,
in which the mark of having no sexual organs was repeatedly adopted
as running through the whole class, although various instances of
sexuality in Cryptogams were known before 1845; Schmidel had described
the sexual organs of the Liverworts about the middle of the previous
century, Hedwig those of the Mosses in 1782, and Vaucher in 1803 had
suggested that the conjugation of Spirogyra among the Algae should be
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