That the leaves are arranged on the stems that produce them according
to fixed geometrical rules had been noticed by Cesalpino and by
Bonnet in the middle of the eighteenth century; but nothing more
resulted than weak attempts at mere description of different cases.
Schimper’s theory is marked by that which is at once its greatest
merit and its fundamental error, the referring of all relations of
position to a single principle. This principle lies in the idea
that growth in a stem has an upward direction in a spiral line, and
that the formation of leaves is a local exaggeration of this spiral
growth. The direction of the spiral line may change in the same
species, or in the same axis, and may even change from leaf to leaf.
The important variations in the arrangement of leaves are not shown
in their longitudinal distances, but in the measure of their lateral
deviations on the stem. The characteristic point in this theory is the
mode of considering these lateral deviations or divergences of the
leaves as they follow one another on an axis, the referring them to a
more general law of position. Means were at the same time skilfully
supplied for discovering the true conditions of arrangement, the
genetic spiral, in cases where the genetic succession of the leaves,
and consequently their divergence, could not be immediately recognised.
After innumerable observations, it appeared that there is a wonderful
variety in the disposition of leaves, but that at the same time a
comparatively small number of these variations commonly occur, and
that these ordinary divergences 1/2, 2/3, 3/8, 8/13, 13/21, etc. have
this remarkable relation to one another, that both the numerator and
denominator of each successive fraction are obtained by adding together
the numerators and denominators of the two preceding fractions, or
the individual fractions named are the successive convergents of a
continuous fraction:—
1
1 + 1
——-
1 + 1
——-
1....
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