His train of thought in search of that which must be called an
individual in the vegetable kingdom is briefly as follows: In forming a
conception of the plant-individual as the unity of a cycle of formation
or a morphological whole, our chief difficulty lies in the division
into parts and the divisibility (Getheiltheit und Theilbarkeit) which
are present in the very different stages of the organic structure of
plants. It is requisite therefore to find the middle way between the
morphological consideration of the individual plant which breaks up
the whole from above downwards, and the physiological which extends it
in the upward direction beyond all limits. Neither the leaf-bearing
shoots, though they are capable of developing into independent plants,
nor the parts of them, which have the same power, neither the single
cells, nor the granules they contain, and least of all the atoms of
dead matter which are the sport of blind forces, would answer to
the idea of the individual in plants. We have therefore to decide
which member of this many-graded series of potences in the cycle of
development subordinated to the species deserves by preference the name
of individual (p. 48). A compromise is then made; it is sufficient to
find a part of the plant which answers above all others to the idea
of the individual, for in this idea there must be two genetic forces,
multiplicity and unity. He then decides for the shoot or bud. ‘In
contemplating the plant-stem which is usually branched, especially
a tree with its many branches, mere instinctive feeling awakens the
suspicion that it is not a single being, a single life, to be classed
with the individual animal or individual man, but that it is a world
of united individuals which spring from one another in a succession of
generations,’ etc. He proceeds to show that this conception, arising as
it does from a sound, natural feeling, is also confirmed by scientific
examination. It appears, however, that many phenomena in the growth
of plants will not fall in well with this instinctive feeling, and
so he says at page 69, ‘We cut the Gordian knot in this way, that
if we have other and sufficient grounds for regarding branches as
individuals, we come to the determination to let every branch pass for
an individual, however strongly the appearance may be against it.’
The shoot is therefore the morphological individual in the plant, and
is analogous to the individual animal. It may certainly be objected,
that we may cut the knot in another way and maintain with Schleiden
that the cells are the individuals in the vegetable kingdom, if we do
not actually arrive by the same path at calling each atom, or at the
other end of the scale the whole self-nourishing plant, an individual,
for about equally strong reasons might be adduced for both one and the
other of these views. It all depends on the point of view we adopt in
such speculations, and on the weight we allow to instinctive feeling
in establishing scientific ideas.
Public-domain text, read in full here on John Shaqi.
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