of alternation of generations, so obvious in the Ferns and Mosses, was
seen in the formation of the seed in the Phanerogams. The changes,
which the alternation of generations passes through from the Muscineae
upwards to the Phanerogams, were, if possible, still more surprising
than the alternation of generations itself.
The reader of Hofmeister’s ‘Vergleichende Untersuchungen’ was presented
with a picture of genetic affinity between Cryptogams and Phanerogams,
which could not be reconciled with the then reigning belief in the
constancy of species. He was invited to recognise a connection
of development which made the most different things appear to be
closely united together, the simplest Moss with Palms, Conifers, and
angiospermous trees, and which was incompatible with the theory of
original types. The assumption that every natural group represents an
idea was here quite out of place; the notion entertained up to that
time of what was really meant by the natural system had to be entirely
altered; it could as little pass for a body of Platonic ideas as for a
mere framework of conceptions. But the effect of the work was great in
respect to the system also; the Cryptogams were now the most important
objects in the study of morphology; the Muscineae were the standard by
which the lower Cryptogams must be tried, the Ferns were the measure
for the Phanerogams. Embryology was the thread which guided the
observer through the labyrinth of comparative and genetic morphology;
metamorphosis now received its true meaning, when every organ could
be referred back to its parent-form, the staminal and carpellary
leaves of the Phanerogams, for example, to the spore-bearing leaves
of the Vascular Cryptogams. That which Häckel, after the appearance
of Darwin’s book, called the phylogenetic method, Hofmeister had
long before actually carried out, and with magnificent success. When
Darwin’s theory was given to the world eight years after Hofmeister’s
investigations, the relations of affinity between the great divisions
of the vegetable kingdom were so well established and so patent, that
the theory of descent had only to accept what genetic morphology had
actually brought to view.
Public-domain text, read in full here on John Shaqi.
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