The literature of mycology based on De Bary’s methods of observation
and its actual results has been enriched by others also in various
directions since 1860; in the case of the Fungi, as in that of the
Algae, it is not possible yet to see to what results investigation
will ultimately lead; but it is one of the fairest fruits of strictly
inductive method, that it has succeeded in smoothing this thorny
and indeed perilous route, where the enquirer is constantly in
danger of being misled, and in satisfying the severest demands of
science. Conclusions have been already reached that are important for
morphology and systematic botany, and among these the establishment
of the nature of the large sporophores, and of processes similar to
the alternation of generations in the higher Cryptogams should be
especially mentioned. But the most important result remains to be
told; it is, that the two classes of Algae and Fungi, hitherto kept
strictly separate, must obviously be now united, and an entirely
new classification adopted, in which Algae and Fungi recur as forms
differing only in habit in various divisions founded on their
morphology[62].
A few words must be given here to the Lichens. They are the division
of the Thallophytes, whose true nature was last recognised, and
that only in modern times; till after 1850 scarcely more was known
of their organisation than Wallroth had discovered in 1825[63],
namely, that green cells, known as gonidia are scattered through the
fungus-like hyphal tissue of the thallus. After Mohl’s investigations
in 1833, it was known that free spores were formed in the tubes of
the fructifications (apothecia), and that a dust collected from the
thallus and consisting of a mixture of gonidia and hyphae was in a
condition to propagate the species. The genetic relation between
the chlorophyll-containing gonidia and the fungus-like hyphae long
continued to be obscure, till at last, after 1868, it was shown that
the gonidia are true Algae, and the hyphal tissue a genuine Fungus, and
that therefore the Lichens are not a class co-ordinating with the Algae
and Fungi, but a division of Ascomycetes, which have this peculiarity,
that they spin their threads round the plants on which they feed, and
take them up into their tissue. De Bary suggested this explanation, but
it was Schwendener who adopted it without reserve and openly declared
it, as much to the surprise as the annoyance of Lichenologists. It may
be foreseen that their opposition will yield to the weight of facts,
which already leave no doubt in the minds of the unprejudiced.
Public-domain text, read in full here on John Shaqi.
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