to fashion what has been seen bit by bit into a clear and connected
representation.
It appears then that progressive improvement of the microscope was
not in itself sufficient to ensure the advance of phytotomy. It would
not indeed be too much to say, that the progress which microscopic
anatomy made step by step with the aid of imperfect instruments
repeatedly gave the impulse to energetic efforts to improve them. Only
practical microscopists could tell where the real defects of existing
instruments lay; it was their anxiety to make them more manageable,
their constant complaints of the poor performance of the optical
part—complaints loudly expressed, especially at the end of the previous
and the beginning of the present century, which urged the opticians
to turn their attention to the microscope and to endeavour to make it
more perfect. Moreover, essential improvements in the instrument were
made by microscopists themselves. Thus Robert Hooke was the first who
in 1760 gave the compound microscope a form convenient for scientific
observation, and Leeuwenhoek developed the powers of the simple
microscope to their highest point. The modern microscope is greatly
indebted for its perfectness to Amici; nor ought the name of von Mohl
to be omitted here, who invented improved methods for microscopic
measurement, and in his work ‘Mikrographie’ (1846) on the construction
of the microscope gave many practical hints to the opticians.
We shall not then make the most important advances in the anatomy of
plants depend as a matter of course and quite passively on the history
of the microscope; they were determined here as in other parts of
botany by a logical necessity of their own; here as elsewhere we have
to fix our eye on the objects pursued by successive enquirers. If for
this purpose we cast a glance over the history of the subject, it
will appear that its founders in the latter half of the seventeenth
century, Malpighi and Grew, were chiefly bent on determining the
connection between the cellular and fibrous elements in the structure
of plants. Two fundamental forms of tissue were assumed from the first,
the succulent cellular tissue composed of chambers or tubes, and, in
contrast to this, the elongated usually fibrous or tubular elementary
organs, the distinction of which into open canals or vessels and
fibres with closed ends continued to be doubtful. The characteristic
feature of this period is, that the investigation of the more delicate
structure is everywhere closely interwoven with reflections on the
function of the elementary organs, and that thus anatomy and physiology
support each other, but not without mutual injury through the
imperfections of both. But the physiological interest far outweighed
the anatomical with the first phytotomists, who used anatomical
research for the purposes of physiology.
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