This is in all essential points Wolff’s theory. We may omit his
statements on the subject of the first formation of leaves at the
growing point and of the development of the parts of the flower, as
well as his physiological views on food and sexuality, which remained
for a long time without influence on the growth of opinion, and mention
only his doctrine of the growth of thickness of the stem. The stem is
originally the prolongation of all the leaf-stalks united together.
As many bundles of vessels are formed in the developed stem as there
are leaves springing from the vegetative axis; each leaf has a single
vascular bundle belonging to it in the stem, in modern phraseology
an inner leaf-trace. The union of these bundles from the different
leaves forms the rind of the stem; but if the leaves are very numerous,
their descending bundles form a closed cylinder, and if the stem is
perennial, the fresh production of leaves every year produces new zones
of wood of this kind every year, which are the yearly rings. This view
of Wolff’s on the growth of the stem in thickness bears an unmistakable
resemblance to the theory afterwards suggested by Du Petit-Thouars,
according to which the roots which descend from the buds are supposed
to effect the thickening of the stem.
The contests between Mirbel and his German antagonists at the beginning
of the present century will bring us back again to the more important
points in Wolff’s theory of the cell. Contemporary botanists paid less
attention to the ‘Theoria Generationis’ than they did to Hedwig’s[71]
phytotomic views, not on the formation of cells, but on the structure
of mature tissue. HEDWIG had given various figures and descriptions of
phytotomic subjects in his ‘Fundamentum Historiae Muscorum’ (1782) and
afterwards in his ‘Theoria Generationis’ (1784); but he treats these
topics at greater length in his treatise ‘De fibrae vegetabilis et
animalis ortu,’ published in 1789, and known to the author of this work
only imperfectly from quotations in later writers. Hedwig’s figures
of histological objects appear to be better than those of any of his
predecessors; they show that he used strong magnifying powers, and that
his glass had a clear field of sight. His defect lay in preconceived
opinions and hasty interpretation of what he observed. In order to
refute Gleichen’s view of the stomata in ferns, he demonstrated the
existence of these organs in many phanerogams, and observed the opening
of the slits, which he named ‘spiracula.’ On the epidermis which he
had removed for the purpose of these observations he saw plainly the
double contour lines bounding the epidermis-cells, and therefore the
cell-walls, which are at right angles to the surface. These he took
for a particular form of vessel, and called them ‘vasa reducentia’ or
‘lymphatica,’ and afterwards ‘vasa exhalantia,’ and he thought that
he had found them again in the interior of parenchymatous tissue,
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