maintains that cells are closed bodies, and proves it well by the
occurrence of cells with coloured juice in the middle of colourless
tissue. Rudolphi represents the orifices of the stomata as encircled
by a roundish rim, which he takes without hesitation for a closing
muscle because the apertures enlarge and diminish. Link is more happy
in taking the part that surrounds the aperture for a cell, or a group
of cells. Rudolphi considers the great cavities in hollow stems and in
the tissue of water-plants as the only air-passages in plants; Link
explains these cavities as gaps caused by the irregular growth of
cellular tissue. With Rudolphi the word vessel means not only vascular
forms in wood, but milk-vessels and resin-ducts also, and to the former
he even transfers Malpighi’s view of the structure of spiral vessels.
Link designates the tubes of the wood only as vessels, combining the
most various forms of them under the term spiral vessels; he excludes
milk-vessels, resin-ducts, and the like from the conception of a
vessel, and in this he is somewhat inconsistent, since he assumes with
Rudolphi that a vessel, in plants as in animals, is a canal for the
conveyance of nutrient sap.
With all these contradictions, the two essays agree in adopting the
old Malpighian view of the growth in thickness of stems, according to
which the new layers of wood are formed from the inner layers of bast,
while between the bast-cells, which are here taken to be identical with
woody fibre, new spiral vessels arise contemporaneously, and, as Link
expressly says, from juices which pour out between the bast-cells.
It is hard to understand how two treatises, so contradictory as
they have been shown to be, could have both received a prize at the
same time, or how the great difference could have been overlooked
between Link’s sensible and well-arranged account of his subject, and
Rudolphi’s uncritical statements, which everywhere rely more on old
authority than on his own observation. It is however certain that
Link’s better production is inferior to Bernhardi’s treatise, unless
we choose to consider the greater copiousness of detail in Link, the
number of his observations, and his acquaintance with the literature
of the subject, as giving him the advantage. His figures, as well as
Rudolphi’s, are not so good as those of Bernhardi.
Public-domain text, read in full here on John Shaqi.
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