But nature has a speedier and more
powerful instrument in the larger pores, which the microscope reveals.
Mirbel does not discuss the question how the fluids are set in motion,
easily disregarding such mechanical difficulties, as was usual in those
days, when vital power was always in reserve to be the moving agent. He
warmly repels the imputation, which Sprengel had made against him, of
having confounded pores and granules, by appealing to his figures; he
says that he has depicted prominences on the outside of the walls of
the dotted vessels, and an orifice in each of them, which his opponents
simply never saw. The question whether these prominences lie on the
inside or the outside of the walls of the vessel has no meaning, if
we suppose with Mirbel that the dividing wall is single; he is only
concerned to enquire whether the perforated projections lie on the one
or on the other side of the wall. He refers Treviranus, who had denied
the presence of the pores, to his description of scalariform vessels,
in which he had himself seen the slits which correspond to the pores.
In comparison with these fundamental questions Mirbel’s further account
of matters of detail does not concern us here. He gave a connected
view of the whole of his doctrine of tissues in the form of aphorisms,
which occupy the second part of his book. Of all that he says on the
five kinds into which he distinguishes vessels the most interesting
is the statement, that diaphragms pierced like a sieve separate the
different members of his ‘beaded’ vessels. We find that the weakest
part of Mirbel’s phytotomy, as of that of his opponents, is his
description of the true vessels (vasa propria), with which he classes
the milk-cells of the Euphorbiae and the resin-ducts of Coniferae, but
he saw clearly enough that the latter were canals inclosed in a layer
of tissue of their own. The third part of the book is devoted to these
forms of tissue, and we learn that he classes not only many kinds of
sieve-cell-bundles, but also true bast-fibres, as those of nettle and
hemp, with his bundles of true vessels. Like his opponents he makes
the growth in thickness in woody stems to be due to transformation of
the inner layer of bast; but he gives a new turn to this view, which
brings it nearer to the modern theory; during the period of vegetation
a delicate tissue with large vessels is developed in Dicotyledons on
the confines of the wood and the bark, and these augment the mass of
the woody body, while a loose cellular tissue is formed on the other
side, destined to replace the constant losses of the outer rind. To
later phytotomists, who understood by the word cambium a thin layer
of tissue constantly engaged in producing wood and rind, Mirbel’s
otherwise indistinct conception of growth in thickness must have become
more indistinct from his using the word cambium not for the layer of
tissue afterwards so called, but for a highly ‘elaborated and purified
Public-domain text, read in full here on John Shaqi.
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