Meyen adopted three fundamental forms of elementary organs in plants;
cells, spiral tubes, and sap-vessels; systems, he says, are formed by
union of similar elementary organs; hence there is a cell-system, a
spiral tube-system, and a system of sap-vessels (vascular system). We
see at once by this classification how closely he follows the ideas
formed before Moldenhawer. The establishment of these three systems is
a retrograde step, since Moldenhawer had already clearly distinguished
between vascular bundles and cell-tissue. Meyen then discusses each
system at length and shows how they are grouped together. He lays
great stress, as he did also at a later period, on the difference in
the characteristic forms of cell-tissue, for which he introduced the
names merenchyma, parenchyma, prosenchyma and pleurenchyma. These
he calls regular cell-tissue, the shapes of the cells being like
geometrical bodies, in opposition to the irregular tissue of Fuci,
Lichens and Fungi. It is a decided improvement on former practice,
and one that marks his later works also, that in connection with the
structure of the solid cell-fabric he discusses the contents of cells
in a special chapter, in which first the matter in solution, then the
granular bodies with organized structure are considered, though with
the latter he classes not only starch-grains, chlorophyll-corpuscles
and the like, but also the spermatozoa in pollen-grains and layers
of thickening matter projecting on the inside of cell-walls, such
as the spiral bands in the elaters of Jungermannieae and several
similar formations. He describes the crystals in vegetable cells at
some length, and finally discusses the movement of the cell-contents
(‘sap’), not omitting that of rotation in the Characeae as observed
by Corti, and in other water-plants. The chapter on intercellular
spaces also shows considerable advance on the views which obtained in
1812; Meyen calls it an account of the spaces produced in cell-tissue
by the union of the cells; the true intercellular passages filled
with air are here distinguished from receptacles of secretions,
resin-passages, gum-passages, oil-passages, and secretion-receptacles
of the nature of cavities. The large air-passages and gaps, such as
occur in water-plants, are a third form of intercellular space; his
air-canals in the wood of oak filled with cell-tissue are obviously
vessels filled with the substance known as thylosis. The form of the
cells in the tissue he thinks is not due to mutual pressure, and he
rejects Kieser’s view that the ideal fundamental form of cells must be
a rhombododecahedron; but he thinks there is a significant resemblance
between the shape of cells and that of basaltic columns.
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