histology began to be more thoroughly studied.
As von Mohl had from the first laid special stress on the peculiar
character of the vascular bundles as compared with other tissue-masses,
so he perceived that the structure of the epidermis and of the
different forms of exterior tissue was thoroughly characteristic, and
he succeeded in arriving at a clearer understanding of the matter
in this case than in the other. Very confused ideas had prevailed
on the subject before he took it up, and we owe to him the best and
most important knowledge which we at present possess. Especially
important were his researches into the origination and true form of
stomata (1838 and 1856), and into the cuticle and its relation to the
epidermis (1842 and 1845). He brought entirely new facts to light
by his study of the development of cork and the outer bark in 1836;
these tissues had scarcely been examined with care till then, and
their formation and relation to the epidermis and the cortical tissue
were quite unknown. In the latter treatise, one of his best, the
difference between the suberous periderm and the true epidermis was
first shown, the various forms of the periderm were described, and the
remarkable fact established that the scaling of the bark was due to
the formation of fine laminae of cork, which, penetrating gradually
into the substance of the cortex, withdraw more and more of it from
its connection with the rest of the living tissue, and as they die off
form by their accumulation a rugged crust, which is the outer bark
surrounding most thick-stemmed trees. The investigation was so thorough
and comprehensive, that later observers, Sanio especially in 1860,
could only add to it some more delicate features in the history of the
process. In the same year appeared his enquiry into the lenticels,
where von Mohl however overlooked what Unger discovered at the same
time (‘Flora,’ 1836), namely, that these forms arise beneath the
stomata; but he at once corrected Unger’s hazardous supposition that
the lenticels are similar forms to the heaps of gemmae on the leaves of
the Jungermannieae. Unger, for his part, was not long in adopting von
Mohl’s explanation of the lenticels as local cork-formations.
Since von Mohl thus distinctly brought out the special character of
the vascular bundles and of the different forms of epidermal tissues,
it must excite surprise that he, like former phytotomists, did not
find himself under the necessity of framing some conception of the
rest of the tissue-masses in their peculiar grouping as a whole, as a
special system, and of classifying and suitably naming the different
forms that compose them, though his examination of Tree-ferns would
seem to have offered him an occasion for doing so. Von Mohl, like his
contemporaries, was satisfied with calling everything that is neither
epidermis, cork or vascular bundle, parenchyma, without distinctly
defining the expression.
Public-domain text, read in full here on John Shaqi.
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