Schleiden’s theory was now definitively set aside, a deeper insight was
obtained into the nature of the cell, and the ideas connected with the
word became broader and more profound. The knowledge of the formation
of cells showed that the cell-walls, which had been hitherto regarded
as the important part, are only secondary products, that the true
living body of the cell is represented by its contents and especially
by the protoplasm. Alexander Braun, relying on numerous researches
into the lower Algae, expressed himself in 1850 (‘Verjüngung,’ p.
244) to the effect that it is an inconvenience that the word cell is
used at one time to designate the cell with its wall, at another time
the cell without its wall, or again the wall without the cell. Since
the contents are the essential part of the cell and form a separate
and individual whole which has its own membrane-like boundary, the
primordial utricle, before the secretion of the membrane of cellulose,
we must either confine the term cell to the enveloping membrane or
to the chamber which it forms and find another name for the body
of the contents, or else call this the true and proper cell. This,
which presents itself at once as the correct mode of conception to
anyone who observes the formation of swarmspores in Algae and Fungi
and many other cases of cell-formation, was from this time forward
a vital point in the doctrine of the cell. Braun contributed also
to the clearing up of the ideas of botanists on this subject by
bringing together under one systematic view and classifying all the
varieties of cell-formation which were known to him up to the year
1850, and especially by a more searching investigation into modes of
conjugation. Henfrey’s contributions (‘Flora’ of 1846 and 1847) rested
entirely on the observations of German botanists, and brought to light
nothing that was independently and essentially new. On the other hand
Hofmeister’s new observations on the development of pollen (1848), and
his many remarks on cell-formation in his epoch-making researches into
embryology in 1851, contributed repeatedly to the deciding of doubtful
points, especially in the behaviour of the nucleus in cell-formation
and the production of the dividing walls. Von Mohl, who in spite
of his own excellent observations maintained up to 1846 a somewhat
undecided attitude of mind in respect to Schleiden’s theory, which was
at that time still in vogue, published in 1851, in his treatise ‘Die
vegetabilische Zelle,’ an excellent summary of the results which had
been so far achieved. In describing cell-division he notices specially
that the new nuclei occupy the centres of the future daughter-cells
before the division of the contents commences; but he still clung to
his old view, that in every instance of cell-division the parting-wall
must form progressively from without inwards, as in Cladophora,
contrary to Nägeli’s and Hofmeister’s correct statements, that cases
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