How little advance had been made towards the true understanding of
the varieties of forms of tissue in the higher plants before 1850 is
shown, for instance, by Schleiden’s account of tissues on page 232 of
his ‘Grundzüge’ of 1845, where parenchyma, intercellular substance,
vessels, vascular bundles, bast-tissue, bast-cells in Apocyneae and
Asclepiadeae, laticiferous vessels, felted tissue, epidermal tissue,
are discussed in this succession in co-ordinated sections of the text.
It is obvious that no well-ordered view of the whole cellular structure
of a plant of the higher order could be obtained in this way. Further
on in the same work, where Schleiden attempts a classification of
vascular bundles, which he distinguishes into closed and open, and
assigns the latter to Dicotyledons, we find the cambium-layer named as
the outer boundary of these open vascular bundles; the bast which lies
outside the cambium was therefore not considered to be a part of the
open vascular bundles, and this necessarily excluded any profitable
comparison of the circumstances in Monocotyledons and Dicotyledons.
The case is still worse in many respects in Schacht’s work already
mentioned, ‘Die Pflanzenzelle’ of 1852, where under the heading ‘Kinds
of vegetable cells’ the histology is discussed in the following
co-ordinated sections; the swarm-filaments of Cryptogams, the spores of
the same, pollen-grains, cells and tissue of Fungi and Lichens, cells
and tissue of Algae, parenchyma and its cells, vessels of the plant,
wood and its cells, bast-cells, stomata, appendicular organs of the
epidermis, cork; then follows a paragraph on the thickening-ring, and
then to the no small astonishment of the reader comes an account of the
vascular bundles, after the vessels, the wood, and the bast-cells have
been already dismissed. That such a mode of presenting the subject is
due to the little insight possessed by the writer into the structure of
the plant as a whole is apparent from simply reading the book, and a
similar confusion of ideas is found in his text-book of 1856.
We find a much better classification of tissues in 1855 in Unger’s
‘Anatomie und Physiologie der Pflanzen’; an account of cells is
followed by a description of cell-complexes, as one of the chief
divisions of the book, and herein of cell-families, cell-tissues, and
cell-fusions. Another chief section is occupied with cell-groups,
and here epidermal formations, air-spaces, sap-receptacles, glands
and vascular bundles are noticed; here certainly the fact has been
overlooked that vascular bundles may be co-ordinated with epidermal
formations, but not air-spaces, sap-receptacles and glands. His last
chief division gives an account of tissue-systems and of the way in
which the vascular bundles are united together in different plants, and
secondary growth in thickness and the activity of the cambium-layer are
described quite in the right connection.
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