way systematic botany became mixed up with theological notions, and
it is easy to understand why the first feeble attempts at a theory of
descent encountered such obstinate, nay, fanatical opposition from
professed systematists, who looked upon the system as something above
nature, a component part of their religion. And if we look back we find
that these views are based on the dogma of the constancy of species,
while Linnaeus’ ‘Philosophia Botanica’ teaches us on what grounds this
dogma rests, where it says, ‘Novas species dari in vegetabilibus
negat generatio continuata, propagatio, observationes quotidianae,
cotyledones.’
In spite of all this one important advance was made by the successors
of Jussieu; the larger groups of genera, the families, were defined
with the certainty and precision, with which Linnaeus had fixed the
boundaries of species and genera, and were supplied with characteristic
marks. They succeeded also in clearly distinguishing various still
larger groups founded on natural affinity, such as the Monocotyledons
and Dicotyledons; the distinction between Cryptogams and Phanerogams
was by degrees better appreciated, though this point could not be
finally settled, so long as it was attempted to reduce the Cryptogams
entirely to the scheme of the Phanerogams. The chief hindrance however
to the advance of systematic botany, at least at the beginning of
this period, lay in the defective morphology enshrined in Linnaeus’
terminology and in his doctrine of metamorphosis. A great improvement
certainly was effected in the early part of the 19th century by De
Candolle’s doctrine of the symmetry of plants,—a doctrine which has
been much undervalued, and that merely on account of its name; it is
really a comparative morphology, and the first serious attempt of the
kind since the time of Jung that has produced any great results; a
series of the most important morphological truths, with which every
botanist is now conversant, were taught for the first time in De
Candolle’s doctrine of symmetry in 1813. But one thing was wanting
not only in Jussieu and De Candolle, but in all the systematists of
this period, with the single exception of Robert Brown, and this
was the history of development. The history of the morphology and
systematic botany of this period shows indeed, that the comparison of
mature forms leads to the recognition of many and highly important
morphological facts; but as long as matured organisms only are
compared, the morphological consideration of them is always disturbed
by the circumstance that the organs to be compared are already adapted
to definite physiological functions, and thus their true morphological
character is often entirely obscured; on the other hand, the younger
the organs are, the less is this difficulty experienced, and this is
the real reason why the history of development is of so great service
to morphology. It was then one of the characteristic features of the
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