It was the XVI century that added to St. Denis' the three tombs of most architectural pretensions, those of Louis XII, Francis I, and Henry II. The monument of Louis XII and Anne of Brittany was undertaken (1516-32) by Jean Juste, who with his brothers had come north from Florence, being among the first to bring into France the ideals of the Renaissance.[34] It has been suggested that the king's and queen's kneeling images are from the studio at Tours of Guillaume Regnault, who for forty years was co-worker with Michel Colombe, last of the great Gothic artists. The priants are still quite French in treatment. Jean Juste made the gisants and his brother and nephew aided with the lesser sculpture. It was Louis XII who ordered artists at Genoa to make, in 1502, the Carrara marble tomb of his father, the poet-duke, Charles d'Orléans, and of his grandfather, the murdered duke of Orléans, builder of Pierrefonds Castle, and son of the art-loving Valois king, Charles V. The tomb of Francis I (1549-59) was designed by Philibert de Lorme. Pierre Bontemps fashioned the bas-reliefs that celebrate the wars in Italy; he and other masters made the _priants_ and _gisants_. The tomb of Henry II and Catherine de Medici (1570) of less artistic value, has a complicated history. The Italian, Primatici, directed the works; Domenico Florentino made the king's kneeling figure, and Germain Pilon his _gisant_; Jerome della Robbia chiseled the queen's death image. To sum up: there are in St. Denis' abbatial three totally different parts, built in different periods. There is Suger's forechurch, in which linger Romanesque echoes; there is the ambulatory of purest Primary Gothic built a little later by the same great abbot; and finally there are nave, transept, and the main parts of the choir erected during the reign of St. Louis in the zenith of Gothic art.