The case of the rumpless fowl is important in relation to the hypothesis of inhibitors. Either tail-production depends on a special factor _TT_, which is diluted, as _Tt_, in the heterozygote; or else there is a tail inhibitor, _II_, which is diluted, as _Ii_, in the heterozygote. In F2 we expect, on the one hypothesis, 25 per cent _tt_, giving no tail, and 25 per cent _TT_, giving tail; on the other hypothesis 25 per cent _ii_, giving tail, and 25 per cent _II_, giving no tail. Actually we get all tailed in some cases; in others 25 per cent with no tail. Which hypothesis best fits the facts? Which is the more probable--that the 25 per cent recessive no-tail should produce a tail (as it were, out of nothing) or that the 25 per cent dominant tail inhibitor should be ineffective, permitting the development of a tail? It is clear that the ontogenetic failure of an inhibitor is easier to understand than the development of a character that is not represented at all in the germ-plasm. This matter is treated in another connection in the next section. But the present point is that it is equally in accord with the facts to regard heredity as initiating and inhibiting processes. If, indeed, processes were not regularly inhibited, they must, when once started, go on indefinitely, as do the hairs of Angora goats and wonder-horses. As we have seen, ontogeny is not completed at hatching or birth. Many characters are at that time undeveloped. Hence, not infrequently the recessive condition is at first seen and is only later replaced by the dominant condition. The reverse sequence will rarely be followed, because development rarely, except in cases of degeneration, moves backward. One of the familiar cases of this sort is human hair-color. In youth this is frequently flaxen, later it becomes light brown, and eventually it may become dark brown. Darwin gives a number of examples in his Chapter XII of Animals and Plants under Domestication. To these I may add some from my own experience. The hybrids between white and gray Java sparrows are at first light and later become of a slaty gray like the dark parent. Many black fowl gain white feathers as they grow older, and every fancier knows that birds with complex white-and-black patterns can usually be "exhibited" only once, on account of loss of "standard" coloration late in life. In these cases the advanced condition in the series of melanic colors appears only late in ontogeny.[13] Similarly Lang (1908, p. 54) finds that in snail hybrids often the young shells have the recessive yellow color, only later in life showing the dominant red color. This is, of course, no reversal of dominance in ontogeny, but mere ontogenesis of pigmentation. So in general, since the recessive condition is absence of the character or its low stage of development and the dominant condition is presence of the full character, the individual in ontogenesis may exhibit in succession