It must not be supposed from these facts that the grains of
chlorophyll, as they exist in living plants, cannot be attacked by the
secretion; for these grains consist of protoplasm merely coloured by
chlorophyll. My son Francis placed a thin slice of spinach leaf,
moistened with saliva, on a leaf of Drosera, and other slices on damp
cotton-wool, all exposed to the same temperature. After 19 hrs. the
slice on the leaf of Drosera was bathed in much secretion from the
inflected tentacles, and was now examined under the microscope. No
perfect grains of chlorophyll could be distinguished; some were
shrunken, of a yellowish-green colour, and collected in the middle of
the cells; others were disintegrated and formed a yellowish mass,
likewise in the middle of the cells. On the other hand, in the slices
surrounded by damp cotton-wool, the grains of chlorophyll were green
and as perfect as ever. My son also placed some slices in artificial
gastric juice, and these were acted on in nearly the same manner as by
the secretion. We have seen that bits of fresh cabbage and spinach
leaves cause the tentacles to be inflected and the glands to pour forth
much acid secretion; and there can be little doubt that it is the
protoplasm forming the grains of chlorophyll, as well as that lining
the walls of the cells, which excites the leaves.
Fat and Oil.—Cubes of almost pure uncooked fat, placed on several
leaves, did not have their angles in the least rounded. We have also
seen that the oil-globules in milk are not digested. Nor does olive oil
dropped on the discs of leaves cause any inflection; but when they are
immersed in olive oil, they become strongly inflected; but to this
subject I shall have to recur. Oily substances are not digested by the
gastric juice of animals.
Starch.—Rather large bits of dry starch caused well-marked inflection,
and the leaves did not re-expand until the fourth day; but I have no
doubt that this was due to the prolonged irritation of the glands, as
the starch continued to absorb the secretion. The particles were not in
the least reduced in size; [page 127] and we know that leaves immersed
in an emulsion of starch are not at all affected. I need hardly say
that starch is not digested by the gastric juice of animals.
Action of the Secretion on Living Seeds.
The results of some experiments on living seeds, selected by hazard,
may here be given, though they bear only indirectly on our present
subject of digestion.
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