It follows from these few facts that different kinds of seeds excite
the leaves in very different degrees; whether this is solely due to the
nature of their coats is not clear. In the case of the cress seeds, the
partial removal of the layer of mucus hastened the inflection of the
tentacles. Whenever the leaves remain inflected during several days
over seeds, it is clear that they absorb some matter from them. That
the secretion penetrates their coats is also evident from the large
proportion of cabbage, raddish, and cress seeds which were killed, and
from several of the seedlings being greatly injured. This injury to the
seeds and seedlings may, however, be due solely to the acid of the
secretion, and not to any process of digestion; for Mr. Traherne
Moggridge has shown that very weak acids of the acetic series are
highly injurious to seeds. It never occurred to me to observe whether
seeds are often blown on to the viscid leaves of plants growing in a
state of nature; but this can hardly fail sometimes to occur, as we
shall hereafter see in the case of Pinguicula. If so, Drosera will
profit to a slight degree by absorbing matter from such seeds.]
A Summary and Concluding Remarks on the Digestive Power of Drosera.
When the glands on the disc are excited either by the absorption of
nitrogenous matter or by mechanical irritation, their secretion
increases in quantity and becomes acid. They likewise transmit [page
129] some influence to the glands of the exterior tentacles, causing
them to secrete more copiously; and their secretion likewise becomes
acid. With animals, according to Schiff,* mechanical irritation excites
the glands of the stomach to secrete an acid, but not pepsin. Now, I
have every reason to believe (though the fact is not fully
established), that although the glands of Drosera are continually
secreting viscid fluid to replace that lost by evaporation, yet they do
not secrete the ferment proper for digestion when mechanically
irritated, but only after absorbing certain matter, probably of a
nitrogenous nature. I infer that this is the case, as the secretion
from a large number of leaves which had been irritated by particles of
glass placed on their discs did not digest albumen; and more especially
from the analogy of Dionaea and Nepenthes. In like manner, the glands
of the stomach of animals secrete pepsin, as Schiff asserts, only after
they have absorbed certain soluble substances, which he designates as
peptogenes. There is, therefore, a remarkable parallelism between the
glands of Drosera and those of the stomach in the secretion of their
proper acid and ferment.
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