In these experiments, some of the short tentacles on the disc, which
would have been directed to the centre, had the leaf been immersed in
an exciting fluid, were now inflected in an exactly opposite direction,
viz. towards the circumference. These tentacles, therefore, had
deviated as much as 180o from the direction which they would have
assumed if their own glands had been stimulated, and which may be
considered as the normal one. Between this, the greatest possible and
no deviation from the normal direction, every degree could be observed
in the tentacles on these several leaves. Notwithstanding the precision
with which the tentacles generally were directed, those near the
circumference of one leaf were not accurately directed towards some
phosphate of lime at a rather distant point on the opposite side of the
disc. It appeared as if the motor [page 246] impulse in passing
transversely across nearly the whole width of the disc had departed
somewhat from a true course. This accords with what we have already
seen of the impulse travelling less readily in a transverse than in a
longitudinal direction. In some other cases, the exterior tentacles did
not seem capable of such accurate movement as the shorter and more
central ones.
Nothing could be more striking than the appearance of the above four
leaves, each with their tentacles pointing truly to the two little
masses of the phosphate on their discs. We might imagine that we were
looking at a lowly organised animal seizing prey with its arms. In the
case of Drosera the explanation of this accurate power of movement, no
doubt, lies in the motor impulse radiating in all directions, and
whichever side of a tentacle it first strikes, that side contracts, and
the tentacle consequently bends towards the point of excitement. The
pedicels of the tentacles are flattened, or elliptic in section. Near
the bases of the short central tentacles, the flattened or broad face
is formed of about five longitudinal rows of cells; in the outer
tentacles of the disc it consists of about six or seven rows; and in
the extreme marginal tentacles of above a dozen rows. As the flattened
bases are thus formed of only a few rows of cells, the precision of the
movements of the tentacles is the more remarkable; for when the motor
impulse strikes the base of a tentacle in a very oblique direction
relatively to its broad face, scarcely more than one or two cells
towards one end can be affected at first, and the contraction of these
cells must draw the whole tentacle into the proper direction. It is,
perhaps, owing to the exterior pedicels being much flattened that they
do not bend quite so accurately to the point of excitement as the [page
247] more central ones. The properly directed movement of the tentacles
is not an unique case in the vegetable kingdom, for the tendrils of
many plants curve towards the side which is touched; but the case of
Drosera is far more interesting, as here the tentacles are not directly
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