In the tenth chapter it was shown that the sensitive- [page 275] ness
of the leaves appears to be wholly confined to the glands and to the
immediately underlying cells. It was further shown that the motor
impulse and other forces or influences, proceeding from the glands when
excited, pass through the cellular tissue, and not along the
fibro-vascular bundles. A gland sends its motor impulse with great
rapidity down the pedicel of the same tentacle to the basal part which
alone bends. The impulse, then passing onwards, spreads on all sides to
the surrounding tentacles, first affecting those which stand nearest
and then those farther off. But by being thus spread out, and from the
cells of the disc not being so much elongated as those of the
tentacles, it loses force, and here travels much more slowly than down
the pedicels. Owing also to the direction and form of the cells, it
passes with greater ease and celerity in a longitudinal than in a
transverse line across the disc. The impulse proceeding from the glands
of the extreme marginal tentacles does not seem to have force enough to
affect the adjoining tentacles; and this may be in part due to their
length. The impulse from the glands of the next few inner rows spreads
chiefly to the tentacles on each side and towards the centre of the
leaf; but that proceeding from the glands of the shorter tentacles on
the disc radiates almost equally on all sides.
When a gland is strongly excited by the quantity or quality of the
substance placed on it, the motor impulse travels farther than from one
slightly excited; and if several glands are simultaneously excited, the
impulses from all unite and spread still farther. As soon as a gland is
excited, it discharges an impulse which extends to a considerable
distance; but afterwards, whilst the gland is secreting and absorbing,
the impulse suffices only to keep the same tentacle [page 276]
inflected; though the inflection may last for many days.
Public-domain text, read in full here on John Shaqi.
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