Concluding Remarks.—The leaves of the three foregoing closely allied
species or varieties are manifestly adapted for catching living
creatures. With respect to the functions of the several parts, there
can be little doubt that the long jointed hairs are sensitive, like
those of Dionaea, and that, when touched, they cause the lobes to
close. That the glands secrete a true digestive fluid and afterwards
absorb the digested matter, is highly probable from the analogy of
Dionaea,—from the limpid fluid within their cells being aggregated into
spherical masses, after they had absorbed an infusion of raw meat,—from
their opaque and granular condition in the leaf, which had enclosed a
beetle for a long time,—and from the clean condition of the integuments
of this insect, as well as of crustaceans (as described by Cohn), which
have been long captured. Again, from the effect produced on the
quadrifid processes by an immersion for 24 hrs. in a solution of
urea,—from the presence of brown granular matter within the quadrifids
of the leaf in which the beetle had been caught,—and from the analogy
of Utricularia,—it is probable that these processes absorb
excrementitious and decaying animal matter. It is a more curious fact
that the points on [page 331] the infolded margins apparently serve to
absorb decayed animal matter in the same manner as the quadrifids. We
can thus understand the meaning of the infolded margins of the lobes
furnished with delicate points directed inwards, and of the broad,
flat, outer portions, bearing quadrifid processes; for these surfaces
must be liable to be irrigated by foul water flowing from the concavity
of the leaf when it contains dead animals. This would follow from
various causes,—from the gradual contraction of the concavity,—from
fluid in excess being secreted,—and from the generation of bubbles of
air. More observations are requisite on this head; but if this view is
correct, we have the remarkable case of different parts of the same
leaf serving for very different purposes—one part for true digestion,
and another for the absorption of decayed animal matter. We can thus
also understand how, by the gradual loss of either power, a plant might
be gradually adapted for the one function to the exclusion of the
other; and it will hereafter be shown that two genera, namely
Pinguicula and Utricularia, belonging to the same family, have been
adapted for these two different functions. [page 332]
CHAPTER XV.
DROSOPHYLLUM—RORIDULA—BYBLIS—GLANDULAR HAIRS OF OTHER PLANTS—CONCLUDING
REMARKS ON THE DROSERACEÆ.
Drosophyllum—Structure of leaves—Nature of the secretion—Manner of
catching insects— Power of absorption—Digestion of animal
substances—Summary on Drosophyllum—Roridula—Byblis—Glandular hairs of
other plants, their power of absorption—Saxifraga—Primula—
Pelargonium—Erica—Mirabilis—Nicotiana—Summary on glandular
hairs—Concluding remarks on the Droseraceae.
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