** Dr. Warming ‘Sur la Diffrence entres les Trichomes,’ Copenhague,
1873, p. 6. ‘Extrait des Videnskabelige Meddelelser de la Soc. d’Hist.
nat. de Copenhague,’ Nos. 10-12, 1872. [page 360]
power was transferred to the basal parts of the hairs, or that the
surface of the leaf has been prolonged upwards at numerous points, so
as to unite with the hairs, thus forming the bases of the inner
tentacles.
The above named three genera, namely Drosophyllum, Roridula, and
Byblis, which appear to have retained a primordial condition, still
bear glandular hairs on both surfaces of their leaves; but those on the
lower surface have since disappeared in the more highly developed
genera, with the partial exception of one species, Drosera binata. The
small sessile glands have also disappeared in some of the genera, being
replaced in Roridula by hairs, and in most species of Drosera by
absorbent papillae. Drosera binata, with its linear and bifurcating
leaves, is in an intermediate condition. It still bears some sessile
glands on both surfaces of the leaves, and on the lower surface a few
irregularly placed tentacles, which are incapable of movement. A
further slight change would convert the linear leaves of this latter
species into the oblong leaves of Drosera anglica, and these might
easily pass into orbicular ones with footstalks, like those of Drosera
rotundifolia. The footstalks of this latter species bear multicellular
hairs, which we have good reason to believe represent aborted
tentacles.
The parent form of Dionaea and Aldrovanda seems to have been closely
allied to Drosera, and to have had rounded leaves, supported on
distinct footstalks, and furnished with tentacles all round the
circumference, with other tentacles and sessile glands on the upper
surface. I think so because the marginal spikes of Dionaea apparently
represent the extreme marginal tentacles of Drosera, the six (sometimes
eight) sensitive filaments on the upper surface, as well as the more
numerous ones in Aldrovanda, representing the central [page 361]
tentacles of Drosera, with their glands aborted, but their
sensitiveness retained. Under this point of view we should bear in mind
that the summits of the tentacles of Drosera, close beneath the glands,
are sensitive.
The three most remarkable characters possessed by the several members
of the Droseraceae consist in the leaves of some having the power of
movement when excited, in their glands secreting a fluid which digests
animal matter, and in their absorption of the digested matter. Can any
light be thrown on the steps by which these remarkable powers were
gradually acquired?
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