Experiment 9.—A chain of fibres of roast meat, as thin as bristles and
moistened with saliva, were placed down one whole side, close to the
narrow, naturally incurved edge of a leaf. In 3 hrs. this side was
greatly incurved along its whole length, and after 8 hrs. formed a
cylinder, about 1/20 of an inch (1.27 mm) in diameter, quite concealing
the meat. This cylinder remained closed for 32 hrs., but after 48 hrs.
was half unfolded, and in 72 hrs. was as open as the opposite margin
where no meat had been placed. As the thin fibres of meat were
completely overlapped by the margin, they were not pushed at all
inwards, across the blade.
Experiment 10.—Six cabbage seeds, soaked for a night in water, were
placed in a row close to the narrow incurved edge of a leaf. We shall
hereafter see that these seeds yield soluble matter to the glands. In 2
hrs. 25 m. the margin was decidedly inflected; in 4 hrs. it extended
over the seeds for about half their breadth, and in 7 hrs. over
three-fourths of their breadth, forming a cylinder not quite closed
along the inner side, and about .7 of an inch (1.778 mm.) in diameter.
After 24 hrs. the inflection had not increased, perhaps had decreased.
The glands which had been brought into contact with the upper surfaces
of the seeds were now secreting freely. In 36 hrs. from the time when
the seeds were put on the leaf the margin had greatly, and after 48
hrs. had completely, re-expanded. As the seeds were no longer held by
the inflected margin, and as the secretion was beginning to fail, they
rolled some way down the marginal channel.
Experiment 11.—Fragments of glass were placed on the margins of two
fine young leaves. After 2 hrs. 30 m. the margin of one certainly
became slightly incurved; but the inflection never increased, and
disappeared in 16 hrs. 30 m. from the time when the fragments were
first applied. With the second leaf there was a trace of incurvation in
2 hrs. 15 m., which became decided in 4 hrs. 30 m., and still more
strongly pronounced in 7 hrs., but after 19 hrs. 30 m. had plainly
decreased. The fragments excited at most a slight and doubtful increase
of the secretion; and in two other trials, no increase could be
perceived. Bits of coal-cinders, placed on a leaf, produced no effect,
either owing to their lightness or to the leaf being torpid.
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