Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
_Date of Last Glacial Epoch and its Bearing on the Measurement of
Geological Time._--Directly we go back from this stable period we come upon
changes both in the forms and in the distribution of species; and when we
pass beyond the last glacial epoch into the Pliocene period we find
ourselves in a comparatively new world, surrounded by a considerable number
of species altogether different from any which now exist, together with
many others which, though still living, now inhabit distant regions. It
seems not improbable that what is termed the Pliocene period, was really
the coming on of the glacial epoch, and this is the opinion of Professor
Jules Marcou.[49] According to our views, a considerable amount of
geographical change must have occurred at the change from the Miocene to
the Pliocene, favouring the refrigeration of the northern hemisphere, and
leading, in the way already pointed out, to the glacial epoch whenever a
high degree of excentricity {123} prevailed. As many reasons combine to
make us fix the height of the glacial epoch at the period of high
excentricity which occurred 200,000 years back, and as the Pliocene period
was probably not of long duration, we must suppose the next great phase of
very high excentricity (850,000 years ago) to fall within the Miocene
epoch. Dr. Croll believes that this must have produced a glacial period,
but we have shown strong reasons for believing that, in concurrence with
favourable geographical conditions, it led to uninterrupted warm climates
in the temperate and northern zones. This, however, did not prevent the
occurrence of local glaciation wherever other conditions led to its
initiation, and the most powerful of such conditions is a great extent of
high land. Now we know that the Alps acquired a considerable part of their
elevation during the latter part of the Miocene period, since Miocene rocks
occur at an elevation of over 6,000 feet, while Eocene beds occur at nearly
10,000 feet. But since that time there has been a vast amount of
denudation, so that these rocks may have been at first raised much higher
than we now find them, and thus a considerable portion of the Alps may have
been more elevated than they are now. This would certainly lead to an
enormous accumulation of snow, which would be increased when the
excentricity reached a maximum, as already fully explained, and may then
have caused glaciers to descend into the adjacent sea, carrying those
enormous masses of rock which are buried in the Upper Miocene of the
Superga in Northern Italy. An earlier epoch of great altitude in the Alps
coinciding with the very high excentricity 2,500,000 years ago, may have
caused the local glaciation of the Middle Eocene period when the enormous
erratics of the Flysch conglomerate were deposited in the inland seas of
Northern Switzerland, the Carpathians, and the Apennines. This is quite in
harmony with the indications of an uninterrupted warm climate and rich
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