Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
_Flora of the Galapagos._--The plants of these islands are so much more
numerous than the known animals, even including the insects, they have been
so carefully studied by eminent botanists, and their relations throw so
much light on the past history of the group, that no apology is needed for
giving a brief outline of the peculiarities and affinities of the flora.
The statements we shall make on this subject will be taken from the Memoir
of Sir Joseph Hooker in the _Linnaean Transactions_ for 1851, founded on
Mr. Darwin's collections, and a later paper by N. J. Andersson in the
_Linnaea_ of 1861, embodying more recent discoveries. {288}
The total number of flowering plants known at the latter date was 332, of
which 174 were peculiar to the islands, while 158 were common to other
countries.[64] Of these latter about twenty have been introduced by man,
while the remainder are all natives of some part of America, though about a
third part are species of wide range extending into both hemispheres. Of
those confined to America, forty-two are found in both the northern and
southern continents, twenty-one are confined to South America, while twenty
are found only in North America, the West Indies, or Mexico. This equality
of North American and South American species in the Galapagos is a fact of
great significance in connection with the observation of Sir Joseph Hooker
that the _peculiar_ species are allied to the plants of temperate America
or to those of the high Andes, while the non-peculiar species are mostly
such as inhabit the hotter regions of the tropics near the level of the
sea. He also observes that the seeds of this latter class of Galapagos
plants often have special means of transport, or belong to groups whose
seeds are known to stand long voyages and to possess great vitality. Mr.
Bentham also, in his elaborate account of the Compositae,[65] remarks on
the decided Central American or Mexican affinities of the Galapagos
species, so that we may consider this to be a thoroughly well-established
fact.
The most prevalent families of plants in the Galapagos are the Compositae
(40 sp.), Gramineae (32 sp.), Leguminosae (30 sp.), and Euphorbiaceae (29
sp.). Of the Compositae most of the species, except such as are common
weeds or shore plants, are peculiar, but there are only two peculiar
genera, allied to Mexican forms and not very distinct; while the genus
Lipochaeta, represented here by a single species, is only found elsewhere
in the Sandwich Islands though it has American affinities.
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