Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
On the whole, the affinities of these birds are, as might be expected,
chiefly with Australia and the Pacific Islands; but they exhibit in the
buzzard, one of the owls, and perhaps in some of the Drepanididae, slight
indications of very rare or very remote communication with America. The
amount of speciality is, however, wonderful, far exceeding that of any
other islands; the only approach to it being made by New Zealand and
Madagascar, which have a much more varied bird fauna and a smaller
_proportionate_ number of peculiar genera. The Galapagos, among the true
oceanic islands, while presenting many peculiarities have only four out of
the ten genera of Passeres peculiar. These facts undoubtedly indicate an
immense antiquity for this group of islands, or the vicinity of some very
ancient land (now submerged), from which some portion of their peculiar
fauna might be derived. For further details as to the affinities and
geographical distribution of the genera and species, the reader must
consult Mr. Scott Wilson's work _The Birds of the Sandwich Islands_,
already alluded to.
_Reptiles._--The only other vertebrate animals are two lizards. One of
these is a very widespread species, _Ablepharus poecilopleurus_, ranging
from the Pacific Islands to West Africa. The other is said to form a
peculiar genus of geckoes, but both its locality and affinities appear to
be somewhat doubtful.
_Land-shells._--The only other group of animals which has been carefully
studied, and which presents features of especial interest, are the
land-shells. These are very numerous, about thirty genera, and between
three and four hundred species having been described; and it is remarkable
that this single group contains as many species of {317} land-shells as all
the other Polynesian Islands from the Pelew Islands and Samoa to the
Marquesas. All the species are peculiar, and about three-fourths of the
whole belong to peculiar genera, fourteen of which constitute the subfamily
Achatinellinae, entirely confined to this group of islands and constituting
its most distinguishing feature. Thirteen genera (comprising sixty-four
species) are found also in the other Polynesian Islands, but three genera
of Auriculidae (Plecotrema, Pedipes, and Blauneria) are not found in the
Pacific, but inhabit--the former genus Australia, China, Bourbon, and Cuba,
the two latter the West Indian Islands. Another remarkable peculiarity of
these islands is the small number of Operculata, which are represented by
only one genus and five species, while the other Pacific Islands have
twenty genera and 115 species, or more than half the number of the
Inoperculata. This difference is so remarkable that it is worth stating in
a comparative form:--
Inoperculata. Operculata. Auriculidae.
Sandwich Islands 332 5 9
Rest of Pacific Islands 200 115 16
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