Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
We now proceed to examine those islands which are the very reverse of the
"oceanic" class, being fragments of continents or of larger islands from
which they have been separated, by subsidence of the intervening land at a
period which, geologically, must be considered recent. Such islands are
always still connected with their parent land by a shallow sea, usually
indeed not exceeding a hundred fathoms deep; they always possess mammalia
and reptiles either wholly or in large proportion identical with those of
the mainland; while their entire flora and fauna is characterised either by
the total absence or comparative scarcity of those endemic or peculiar
species and genera which are so striking a feature of almost all oceanic
islands. Such islands will, of course, differ from each {332} other in
size, in antiquity, and in the richness of their respective faunas, as well
as in their distance from the parent land and the facilities for
intercommunication with it; and these diversities of conditions will
manifest themselves in the greater or less amount of speciality of their
animal productions.
This speciality, when it exists, may have been brought about in two ways. A
species or even a genus may on a continent have had a very limited area of
distribution, and this area may be wholly or almost wholly contained in the
separated portion or island, to which it will henceforth be peculiar. Even
when the area occupied by a species is pretty equally divided at the time
of separation between the island and the continent, it may happen that it
will become extinct on the latter, while it may survive on the former,
because the limited number of individuals after division may be unable to
maintain themselves against the severer competition or more contrasted
climate of the continent, while they may flourish, under the more
favourable insular conditions. On the other hand, when a species continues
to exist in both areas, it may on the island be subjected to some
modifications by the altered conditions, and may thus come to present
characters which differentiate it from its continental allies and
constitute it a new species. We shall in the course of our survey meet with
cases illustrative of both these processes.
The best examples of recent continental islands are Great Britain and
Ireland, Japan, Formosa, and the larger Malay Islands, especially Borneo,
Java, and Celebes; and as each of these presents special features of
interest, we will give a short outline of their zoology and past history in
relation to that of the continents from which they have recently been
separated, commencing with our own islands, to which the present chapter
will be devoted.
Public-domain text, read in full here on John Shaqi.
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