Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
Here then we have a group in which there is no question of the means of
dispersal; and where the difficulties that present themselves are not how
the species reached the remote localities in which they are now found, but
rather why they have not established themselves in {370} many other
stations which, so far as we can judge, seem equally suitable to them. Yet
it is a curious fact, that the phenomena of distribution actually presented
by this group do not essentially differ from those presented by the higher
flowering plants which have apparently far less diffusive power, as we
shall find when we come to treat of the floras of oceanic islands; and we
believe that the explanation of this is, that the life of _species_, and
especially of _genera_, is often so prolonged as to extend over whole
cycles of such terrestrial mutations as we have just referred to; and that
thus the majority of plants are afforded means of dispersal which are
usually sufficient to carry them into all suitable localities on the globe.
Hence it follows that their actual existence in such localities depends
mainly upon vigour of constitution and adaptation to conditions just as it
does in the case of the lower and more rapidly diffused groups, and only
partially on superior facilities for diffusion. This important principle
will be used further on to afford a solution of some of the most difficult
problems in the distribution of plant life.[87]
_Concluding Remarks on the Peculiarities of the British Fauna and
Flora._--The facts, now I believe for the first time brought together,
respecting the peculiarities of the British fauna and flora, are sufficient
to show that there is considerable scope for the study of geographical
distribution even in so apparently unpromising a field as one of the most
recent of continental islands. Looking at the general bearing of these
facts, they prove, that the idea so generally entertained as to the
biological identity of the British Isles with the adjacent continent is not
altogether correct. Among birds we have undoubted peculiarities in at least
three instances; peculiar fishes are much more numerous, and in this case
the fact that the Irish species {371} are almost all different from the
British, and those of the Orkneys distinct from those of Scotland, renders
it almost certain that the great majority of the fifteen peculiar British
fishes are really peculiar and will never be found on the European
Continent. The mosses and Hepaticae also have been sufficiently collected
in Europe to render it pretty certain that the more remarkable of the
peculiar British forms are not found there; why therefore, it may be well
asked, should there not be a proportionate number of peculiar British
insects? It is true that numerous species have been first discovered in
Britain, and, subsequently, on the continent; but we have many species
which have been known for twenty, thirty, or forty years, some of which are
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