Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
pervade all the marine deposits formed over a wide area so long as the
state of glaciation continues, we are driven to the conclusion that the
last glacial epoch of the northern hemisphere was exceptional, and was not
preceded by numerous similar glacial epochs throughout Tertiary and
Secondary time.
But although glacial epochs (with the one or two exceptions already
referred to) were certainly absent, considerable changes of climate may
have frequently occurred, and these would lead to important changes in the
organic world. We can hardly doubt that some such change occurred between
the Lower and Upper Cretaceous periods, the floras of which exhibit such an
extraordinary contrast in general character. We have also the testimony of
Mr. J. S. Gardner, who has long worked at the fossil floras of the Tertiary
deposits, and who states, that {93} there is strong negative and some
positive evidence of alternating warmer and colder conditions, not glacial,
contained not only in English Eocene, but all Tertiary beds throughout the
world.[28] In the case of marine faunas it is more difficult to judge, but
the numerous changes in the fossil remains from bed to bed only a few feet
and sometimes a few inches apart, may be sometimes due to change of
climate; and when it is recognised that such changes have probably occurred
at all geological epochs and their effects are systematically searched for,
many peculiarities in the distribution of organisms through the different
members of one deposit may be traced to this cause.
_General View of Geological Climates as dependent on the Physical Features
of the Earth's Surface._--In the preceding chapters I have earnestly
endeavoured to arrive at an explanation of geological climates in the
temperate and Arctic zones, which should be in harmony with the great body
of geological facts now available for their elucidation. If my conclusions
as here set forth diverge considerably from those of Dr. Croll, it is not
from any want of appreciation of his facts and arguments, since for many
years I have upheld and enforced his views to the best of my ability. But a
careful re-examination of the whole question has now convinced me that an
error has been made in estimating the comparative effect of geographical
and astronomical causes on changes of climate, and that, while the latter
have undoubtedly played an important part in bringing about the glacial
epoch, it is to the former that the mild climates of the Arctic regions are
almost entirely due. If I have now succeeded in approaching to a true
solution of this difficult problem, I owe it mainly to the study of Dr.
Croll's writings, since my theory is entirely based on the facts and
principles so clearly set forth in his admirable papers on "Ocean Currents
in relation to the Distribution of Heat over the Globe." The main features
of this theory as distinct from that of Dr. Croll I will now endeavour to
summarise.
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