Island Life; Or, The Phenomena and Causes of Insular Faunas and FlorasWallace, Alfred Russel
Science
Island Life; Or, The Phenomena and Causes of Insular Faunas and Floras
Wallace, Alfred Russel
Biogeography; Glacial epoch; Island ecology
As an indication of the periods usually contemplated by geologists, we may
refer to Sir Charles Lyell's calculation in the tenth edition of his
_Principles of Geology_ (omitted in later editions), by which he arrived at
240 millions of years as having probably elapsed since the Cambrian
period--a very moderate estimate in the opinion of most geologists. This
calculation was founded on the rate of modification of the species of
mollusca; but much more recently Professor Haughton has arrived at nearly
similar figures from a consideration of the rate of formation of rocks and
their known maximum thickness, whence he deduces a maximum of 200 millions
of years for the whole duration of geological time, as indicated by the
series of stratified formations.[30] But in the opinion of all our first
naturalists and geologists, the period occupied in the formation of the
known stratified rocks only represents a portion, and perhaps a small
portion, of geological time. In the sixth edition of the _Origin of
Species_ (p. 286), Mr. Darwin says: "Consequently, if the theory be true,
it is indisputable that before the lowest Cambrian stratum was deposited
long periods elapsed, as long as, or probably far longer than, the whole
interval from the Cambrian age to the present day; and that during these
vast periods the world swarmed with living creatures." Professor Huxley, in
his anniversary address to the Geological Society in 1870, adduced a number
of special cases showing that, on the theory of development, almost all the
higher forms of life must have existed during the Palaeozoic period. Thus,
from the fact that almost the whole of the Tertiary period has been
required to convert the ancestral Orohippus into the true horse, he
believes that, in order to have time for the much greater change of the
ancestral Ungulata into the two great odd-toed and even-toed divisions (of
which change there is no trace even among the earliest Eocene mammals), we
should require a large portion, if not the whole, of the Mesozoic or
Secondary period. Another case is furnished by the bats and whales, both of
which strange modifications of the {101} mammalian type occur perfectly
developed in the Eocene formation. What countless ages back must we then go
for the origin of these groups, the whales from some ancestral carnivorous
animal, and the bats from the insectivora! And even then we have to seek
for the common origin of carnivora, insectivora, ungulata, and marsupials
at a far earlier period; so that, on the lowest estimate, we must place the
origin of the mammalia very far back in Palaeozoic times. Similar evidence
is afforded by reptiles, of which Professor Huxley says: "If the very small
differences which are observable between the crocodiles of the older
Secondary formations and those of the present day furnish any sort of an
approximation towards an estimate of the average rate of change among
reptiles, it is almost appalling to reflect how far back in Palaeozoic
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