"The natural movements of the Medusa precisely resemble those of its
marine congeners. More particularly, these movements resemble those
of the marine species which do not swim continuously, but indulge in
frequent pauses. In water at the temperature of that in the Victoria
lily-house (85° Fahr.), the pauses are frequent, and the rate of
the rhythm irregular, suddenly quickening and suddenly slowing even
during the same bout, which has the effect of giving an almost
intelligent appearance to the movements. This is especially the case
with young specimens. In colder water (65° to 75°) the movements are
more regular and sustained; so that, guided by the analogy furnished
by my experiments on the marine forms, I infer that the temperature
of the natural habitat of this Medusa cannot be so high as that of
the water in the Victoria lily-house. In water of that temperature
the rate of the rhythm is enormously high, sometimes rising to three
pulsations per second. But by progressively cooling the water,
this rate may be progressively lowered, just as in the case of the
marine species; and in water at 65°, the maximum rate that I have
observed is eighty pulsations per minute. As the temperature at which
the greatest activity is displayed by the fresh-water species is a
temperature so high as to be fatal to all the marine species which I
have observed, the effects of cooling are, of course, only parallel
in the two cases when the effects of a series of higher temperatures
in the one case are compared with those of a series of lower
temperatures in the other. Similarly, while a temperature of 70° is
fatal to all the species of marine Medusæ which I have examined,
it is only a temperature of 100° that is fatal to the fresh-water
species. Lastly, while the marine species will endure any degree of
cold without loss of life, such is not the case with the fresh-water
species. Marine Medusæ, after having been frozen solid, will, when
gradually thawed out, again resume their swimming movements; but this
fresh-water Medusa is completely destroyed by freezing. Upon being
thawed out, the animal is seen to have shrunk into a tiny ball, and
it never again recovers either its life or its shape.
"The animal seeks the sunlight. If one end of the tank is shaded, all
the Medusæ congregate at the end which remains unshaded. Moreover,
during the daytime they swim about at the surface of the water; but
when the sun goes down they subside, and can no longer be seen. In
all these habits they resemble many of the sea-water species. They
are themselves non-luminous.
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