[Illustration: Fig. 49.]
But it is in the case of Echinus that these righting movements become
most interesting, from the fact that they are so much more difficult
to accomplish than they are in the case of the Star-fishes. For while
a Star-fish is provided with flat, flexible, and muscular rays,
comprising a small and light mass in relation to the motive power,
an Echinus is a rigid, non-muscular, and globular mass, whose only
motive power available for conducting the manoeuvre is that which is
supplied by its relatively feeble ambulacral feet. It is, therefore,
scarcely surprising that unless the specimens chosen for these
observations are perfectly fresh and vigorous, they are unable to
right themselves at all; they remain permanently inverted till they
die. But if the specimens are fresh and vigorous, they are sooner
or later sure to succeed in righting themselves, and their method
of doing so is always the same. Two, or perhaps three, adjacent
rows of suckers are chosen out of the five, as the rows which are
to accomplish the task (Fig. 49). As many feet upon the rows as can
reach the floor of the tank are protruded downwards and fastened
firmly to the floor; their combined action then serves to tilt the
globe slightly over in their own direction, the anchoring feet on
the other or opposite rows meanwhile releasing their hold of the
tank to admit of this tilting (Fig. 50). The effect of this tilting
is to enable the next feet in the active ambulacral rows to touch
the floor of the tank, and, when they have established their hold,
they assist in increasing the tilt; then the next feet in the series
lay hold, and so on, till the globe slowly but steadily rises upon
its equator (Fig. 51). The difficulty of raising such a heavy mass
into this position by means of the slender motive power available
can be at once appreciated on witnessing the performance, so that
one is surprised, notwithstanding the co-ordination displayed by all
the suckers, that they are able to accomplish the work assigned to
them. That the process is in truth a very laborious one is manifest,
not only from the extreme slowness with which it takes place, but
also because, as already observed, in the case of not perfectly
strong specimens complete failure may attend the efforts to reach the
position of resting on the equator--the Echinus, after rearing up a
certain height, becoming exhausted and again falling back upon its
ab-oral pole. Moreover, in some cases it is interesting to observe
that when the equator position has been reached with difficulty, the
Echinus, as it were, gives itself a breathing space before beginning
the movement of descent--drawing in all its pedicels save those
which hold it securely in the position to which it has attained, and
remaining in a state of absolute quiescence for a prolonged time. It
then suddenly begins to protrude all its feet again, and to continue
its manoeuvre. At any time during such a period of rest, a stimulus
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