If with a sharp scalpel an incision be made through the tenuous
contractile sheet of the sub-umbrella of Aurelia, in a marvellously
short time the injury is repaired. Thus, for instance, if such an
incision be carried across the whole diameter of the sub-umbrella,
so as entirely to divide the excitable tissues into two parts while
the gelatinous tissues are left intact, the result of course is that
physiological continuity is destroyed between the one half of the
animal and the other, while the form of the whole animal remains
unchanged--the much greater thickness of the uninjured gelatinous
tissues serving to preserve the shape of the umbrella. But although
the contractile sheet which lines the umbrella is thus completely
severed throughout its whole diameter, it again reunites, or heals
up, in from four to eight hours after the operation.
CHAPTER V.
EXPERIMENTS IN SECTION OF NAKED-EYED MEDUSÆ.
_Distribution of Nerves in Sarsia._
My experiments have shown that the nervous system in the naked-eyed
Medusæ is more highly organized, or integrated, than it is in the
covered-eyed Medusæ; for whereas in the latter I obtained no evidence
of the gathering together of nerve-fibres into definite bundles or
trunks (the plexus being evenly distributed over the entire surface
of the neuro-muscular sheet lining the umbrella), in the former I
found abundant evidence of this advance in organization. And as the
experiments in this connection serve to substantiate the histological
researches of Professors Haeckel, Schultz, Eimer, and Hertwig, in
as far as the distribution of the main nerve-trunks is concerned, I
shall here detail at some length the character and results of these
experiments in the case of Sarsia.
The occurrence of reflex action in Sarsia is of a very marked and
unmistakable character. I may begin by stating that when any part
of the internal surface of the bell is irritated, the manubrium
responds; but as there is no evidence of ganglia occurring in the
manubrium, this cannot properly be regarded as a case of reflex
action. But now the converse of the above statement is likewise true,
viz. that when any part of the manubrium is irritated, the bell
responds; and it is in this that the unequivocal evidence of reflex
action consists, for while the sympathy of the manubrium with the
bell is not in the least impaired by removing the marginal ganglia of
the latter, the sympathy of the bell with the manubrium is by this
operation entirely destroyed.
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