In the Aurelia of a portion of which the accompanying woodcut (p.
102) is a representation, seven of the lithocysts were removed,
while the remaining one was almost entirely isolated from the
general contractile tissue by the incisions _aa_, _bb_, _cc_. The
lithocyst continued to animate the tissue-area _xxxx_, and through
the connecting passage _y_ the contraction-waves spread over the
remainder of the sub-umbrella tissue _zzzz_. So far, of course, the
facts were normal; but very frequently it was observed that the
contraction-waves did not start from the lithocyst, or from the area
_xxxx_, but from the point o in the area _zz_. After this origination
of the contraction-waves from the point _o_ had been observed a great
number of times, I removed the lithocyst. The effect was not only
to prevent the further origination of contraction-waves in the area
_xxxx_, but also to prevent their further origination from the point
_o_, the entire umbrella thus becoming paralyzed. Hence, before the
removal of the lithocyst, the contraction-waves which originated at
the point _o_, no less than those which originated at the lithocyst
itself, must in some way or other have been due to the ganglionic
influence emanating from the lithocyst and asserting itself at the
distant point _o_.
[Illustration: Fig. 21.]
This property, which lithocysts sometimes present, of asserting their
ganglionic influence at a distance from their own locality, can only,
I think, be explained by supposing that at the point where under
these circumstances the contractions originate, there are situated
some scattered ganglionic cells of considerable functional power,
but yet not of power enough to originate contraction-waves unless
re-enforced by some stimulating influence, which reaches them from
the lithocyst through the nervous plexus.
_Regeneration of Tissues._
The only facts which remain to be stated in the present chapter
have reference to the astonishing rapidity with which the excitable
tissues of the Medusæ regenerate themselves after injury. In this
connection I have mainly experimented on Aurelia aurita, and shall,
therefore, confine my remarks to this one species.
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