Edward Forbes was in the habit of asserting that the similarity of the organic contents of distant formations was 'prima facie' evidence, not of their similarity, but of their difference of age; and holding as he did the doctrine of single specific centres, the conclusion was as legitimate as any other; for the two districts must have been occupied by migration from one of the two, or from an intermediate spot, and the chances against exact coincidence of migration and of imbedding are infinite. In point of fact, however, whether the hypothesis of single or of multiple specific centres be adopted, similarity of organic contents cannot possibly afford any proof of the synchrony of the deposits which contain them; on the contrary, it is demonstrably compatible with the lapse of the most prodigious intervals of time, and with the interposition of vast changes in the organic and inorganic worlds, between the epochs in which such deposits were formed. On what amount of similarity of their faunae is the doctrine of the contemporaneity of the European and of the North American Silurians based? In the last edition of Sir Charles Lyell's 'Elementary Geology' it is stated, on the authority of a former President of this Society, the late Daniel Sharpe, that between 30 and 40 per cent. of the species of Silurian Mollusca are common to both sides of the Atlantic. By way of due allowance for further discovery, let us double the lesser number and suppose that 60 per cent. of the species are common to the North American and the British Silurians. Sixty per cent. of species in common is, then, proof of contemporaneity. Now suppose that, a million or two of years hence, when Britain has made another dip beneath the sea and has come up again, some geologist applies this doctrine, in comparing the strata laid bare by the upheaval of the bottom, say, of St. George's Channel with what may then remain of the Suffolk Crag. Reasoning in the same way, he will at once decide the Suffolk Crag and the St. George's Channel beds to be contemporaneous; although we happen to know that a vast period (even in the geological sense) of time, and physical changes of almost unprecedented extent, separate the two. But if it be a demonstrable fact that strata containing more than 60 or 70 per cent. of species of Mollusca in common, and comparatively close together, may yet be separated by an amount of geological time sufficient to allow of some of the greatest physical changes the world has seen, what becomes of that sort of contemporaneity the sole evidence of which is a similarity of facies, or the identity of half a dozen species, or of a good many genera? And yet there is no better evidence for the contemporaneity assumed by all who adopt the hypothesis of universal faunae and florae, of a universally uniform climate, and of a sensible cooling of the globe during geological time.