_Evernia furfuracea_ with a basal attachment[374], and with a partly
horizontal and partly upright growth, has a dorsiventral thallus, dark
greyish-green above and black beneath, with occasional rhizinae towards
the base. The cortex of both surfaces belongs to the “decomposed” type;
the gonidial zone lies below the upper surface, and the medullary tissue
is of loose hyphae. In certain forms of the species isidia are abundant
on the upper surface, a character of foliose rather than of fruticose
lichens. _E. furfuracea_ grows on trees and very frequently on palings.
[Illustration: Fig. 59. _Evernia prunastri_ Ach. (M. P., _Photo._).]
_E. prunastri_, the second species of the genus, is more distinctly
upright in habit, with a penetrating basal hold-fast and upright
strap-shaped branching fronds, light-greyish green on the “upper” surface
and white on the other (Fig. 59). The internal structure is sub-radiate;
both cortices are “decomposed”; the gonidial zone consists of somewhat
loose groups of algae, very constant below the “upper” surface, with an
occasional group in the pith near to the lower cortex in positions that
are more exposed to light. There is also a tendency for the gonidial zone
to pass round the margin and spread some way along the under side. The
medulla is of loose arachnoid texture and the whole plant is very limp
when moist. It grows on trees, often in dense clusters.
3. FRUTICOSE AND FILAMENTOUS
A. GENERAL STRUCTURE OF THALLUS
The conditions of strain and tension in the upright plant are entirely
different from those in the decumbent thallus, and to meet the new
requirements, new adaptations of structure are provided either in the
cortex or in the medulla.
CORTICAL STRUCTURES. With the exception of the distinctly
plectenchymatous cortex, all the other types already described recur in
fruticose lichens; in various ways they have been modified to provide not
only covering but support to the fronds.
_a._ =The fastigiate cortex.= This reaches its highest development in
_Roccella_ in which the branched hyphal tips, slightly clavate and
thick-walled, lie closely packed in palisade formation at right angles
to the main axis (Fig. 45). They afford not only bending power, but give
great consistency to the fronds. The cortex is further strengthened
in _R. fuciformis_[375] by the compact arrangement of the medullary
hyphae that run parallel with the surface, and among which occur single
thick-walled filaments. The plant grows on maritime rocks in very exposed
situations; and the narrow strap-shaped fronds, as stated above, may
attain a length of 30 cm., though usually they are from 10 to 18 cm. in
height. The same type of cortex, but less highly differentiated, affords
a certain amount of stiffness to the cylindrical much weaker fronds of
_Thamnolia_.
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