C. ALGAE THAT FORM CEPHALODIA
The algae of the cephalodia belong mostly to genera that form the normal
gonidia of other lichens. They are:
_Stigonema_,—in _Lecanora gelida_, _Stereocaulon_, _Pilophorus robustus_,
and _Lecidea pelobotrya_.
_Scytonema_,—a rare constituent of cephalodia.
_Nostoc_,—the most frequent gonidium of cephalodia. It occurs in those of
the genera _Sticta_, _Lobaria_, _Peltigera_, _Nephroma_, _Solorina_ and
_Psoroma_; occasionally in _Stereocaulon_ and in _Lecidea pallida_.
_Lyngbya_ and _Rivularia_,—rarely present, the latter in _Sticta
oregana_[462].
_Chroococcus_ and _Gloeocapsa_,—also very rare.
_Scytonema_, _Chroococcus_, _Gloeocapsa_ and _Lyngbya_ are generally
found in combination with some other cephalodia-building alga, though
Nylander[463] found _Scytonema_ alone in the lobulate cephalodia of
_Sphaerophorus stereocauloides_, a New Zealand lichen, and the only
species of that genus in which cephalodia are developed; and Hue[460]
records _Gloeocapsa_ as forming internal cephalodia in two species of
_Aspicilia_. Bornet[464] found _Lyngbya_ associated with _Scytonema_ in
the cephalodia of _Stereocaulon ramulosum_, and, in the same lichen,
Forssell[465] found, in the several cephalodia of one specimen, _Nostoc_,
_Scytonema_, and _Lyngbya_, while, in those of another, _Scytonema_ and
_Stigonema_ were present. In the latter instance these algae were living
free on the podetium. Forssell[465] also determined two different algae,
_Gloeocapsa magma_ and _Chroococcus turgidus_, present in a cephalodium
on _Lecidea panaeola_ var. _elegans_.
As a general rule only one kind of alga enters into the formation of the
cephalodia of any species or genus. A form of _Nostoc_, for instance,
is invariably the gonidial constituent of these bodies in the genera,
_Lobaria_, _Sticta_, etc. In other lichens different blue-green algae,
as noted above, may occupy the cephalodia even on the same specimen.
Forssell finds alternative algae occurring in the cephalodia of:
_Lecanora gelida_ and _Lecidea illita_ contain either _Stigonema_ or
_Nostoc_;
_Lecidea panaeola_, with _Gloeocapsa_, _Stigonema_ or _Chroococcus_;
_Lecidea pelobotrya_, with _Stigonema_ or _Nostoc_;
_Pilophorus robustus_, with _Gloeocapsa_, _Stigonema_, or _Nostoc_.
[Illustration: Fig. 77. _Lecanora gelida_ Ach. _a_, lobate cephalodia ×
12 (after Zopf).]
Riddle[466] has employed cephalodia with their enclosed algae as
diagnostic characters in the genus _Stereocaulon_. When the alga is
_Stigonema_, as in _S. paschale_, etc., the cephalodia are generally
very conspicuous, grey in colour, spherical, wrinkled or folded, though
sometimes black and fibrillose (_S. denudatum_). Those containing
_Nostoc_ are, on the contrary, minute and are coloured verdigris-green
(_S. tomentosum_ and _S. alpinum_).
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