In the earliest stages, according to Linkola, a small group of algae may
be observed in the cortical tissue of the _Peltigera_ apart from the
gonidial zone and near the upper surface. Gradually a protruding head
is formed which is at first covered over with a brown cortical layer
one cell thick. The head increases and becomes more lobate in form,
being attached to the thallus at the base by a very narrow neck and
more loosely at other parts of the scale. In older scales, the gonidia
are entirely separated from those of the thallus, and a dark-brown
cortex several cells in thickness covers over the top and sides; there
is a colourless layer of plectenchyma beneath. At this advanced stage
the scales are almost completely superficial and correspond with the
cephaloidal rather than with the isidial type of formation. The algae
even in the very early stages are distinct from the gonidial zone and the
whole development, if isidial, must be considered as somewhat abnormal.
3. SOREDIA
A. STRUCTURE AND ORIGIN OF SOREDIA
Soredia are minute separable parts of the lichen thallus, and are
composed of one or more gonidia which are clasped and surrounded by
the lichen hyphae (Fig. 80). They occur on the surface or margins of
the thallus of a fairly large number of lichens either in a powdery
excrescence or in a pustule-like body comprehensively termed a “soralium”
(Fig. 81). The soralia vary in form and dimensions according to the
species. Each individual soredium is capable of developing into a new
plant; it is a form of vegetative reproduction characteristic of lichens.
[Illustration: Fig. 80. Soredia. _a_, of _Physcia pulverulenta_ Nyl.;
_b_, of _Ramalina farinacea_ Ach. × 600.]
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