These results tally well with those obtained by M. and Mme Moreau[616],
though they make no mention of any trichogyne. They found that the
terminal cells of the ascogenous hyphae were transformed into asci, and
the two nuclei in these cells fused—the only fusion that took place. In
_Nephromium_, one of the same family, the case for apogamy is not so
clear; but Fünfstück found no trichogynes, and though spermogonia were
present on the thallus, they were always somewhat imperfectly developed.
Sturgis[617] supplemented these results in his study of other lichens
containing blue-green algae. In species of _Heppia_, _Pannaria_,
_Hydrothyria_, _Stictina_ and _Ricasolia_, he failed to find any evidence
of fertilization by spermatia.
_Solorina_, also a member of Peltigeraceae, was added to the list of
apogamous genera by Metzger[618] and his work was confirmed and amplified
by Baur[619]: certain hyphae of the gonidial zone branch out into
larger ascogonial cells which increase by active intercalary growth, by
division and by branching, and so gradually give rise to the ascogenous
hyphae and finally to the asci. Baur looked on this and other similar
formations as instances of degeneration from the normal carpogonial
type of development. Moreau[620] (Fernand and Mme) have also examined
_Solorina_ with much the same results: the paraphyses rise first from
cells that have been produced by the gonidial hyphae; later, ascogenous
hyphae are formed and spread horizontally at the base of the paraphyses,
finally giving rise at their tips to the asci. Metzger[618] had further
discovered that spermogonia were absent and trichogynes undeveloped
in two very different crustaceous lichens, _Acarospora_ (_Lecanora_)
_glaucocarpa_ and _Verrucaria calciseda_, the latter a pyrenocarpous
species and, as the name implies, found only on limestone.
Public-domain text, read in full here on John Shaqi.
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