They tend to congregate on, and indeed are practically restricted to
the better lighted portions of the thallus. On the fronds of foliose
forms, they appear, for instance, on the swollen pustules of _Umbilicaria
pustulata_, while in _Lobaria pulmonaria_, they are mostly lodged in
the ridges that surround the depressions in the thallus. In _Parmelia
conspersa_, _Urceolaria_ (_Diploschistes_) _scruposa_ and some others,
they occasionally invade the margins of the apothecium or even the
apothecial disc as in _Lichina_. Forssell[712] found that a spermogonium
had developed among cells of _Gloeocapsa_ that covered the disc of a
spent apothecium of _Pyrenopsis haematopis_.
In fruticose lichens such as _Usnea_, _Ramalina_, etc. they occur near
the apex of the fronds, and in _Cladonia_ they occupy the tips of the
ascyphous podetia or the margins of the scyphi. In some _Cladoniae_,
however, spermogonia are produced on the basal squamules, more rarely on
the squamules that clothe the podetia.
_b._ FORM AND SIZE. Spermogonia are specifically constant in form, the
same type being found on the same lichen species all over the globe. The
larger number are entirely immersed and are ovoid or roundish (Fig. 111 A)
or occasionally somewhat flattened bodies (_Nephromium laevigatum_), or
again, but more rarely, they are irregular in outline with an infolding
of the walls that gives the interior a chambered form (Fig. 111 B)
(_Lichina pygmaea_); but all of these are only visible as minute points
on the thallus.
[Illustration: Fig. 111. Immersed spermogonia. A, globose in _Parmelia
acetabulum_ Dub. × 600; B, with infolded walls in _Lecidea_ (_Psora_)
_testacea_ Ach. × 144 (after Glück).]
A second series, also immersed, are borne in small protuberances of the
thallus. These very prominent forms are rarely found in crustaceous
lichens, but they are characteristic of such well-known species as
_Ramalina fraxinea_, _Xanthoria parietina_, _Ricasolia amplissima_,
_Baeomyces roseus_, etc. Other spermogonia project slightly above the
level of the thallus, as in _Cladonia papillaria_ and _Lecidea lurida_;
while in a few instances they are practically free, these last strikingly
exemplified in _Cetraria islandica_ where they occupy the small
projections or cilia (Fig. 112) that fringe the margins of the lobes;
they are free also in most species of _Cladonia_.
In size they vary from such minute bodies as those in _Parmelia
exasperata_ which measure 25-35 µ in diam., up to nearly 1 mm. in Lobaria
_laetevirens_. As a rule, they range from about 150 µ to 400 µ across
the widest part, and are generally rather longer than broad. They open
above by a small slit or pore called the ostiole about 20 µ to 100 µ wide
which is frequently dark in colour. In one instance, in _Icmadophila
aeruginosa_, Nienburg[713] has described a spermogonium with a wide
opening, the spermatiophores being massed in palisade formation along the
bottom of a cup-like structure.
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