3. The _Cladonia_ type: a branching sporophore, each branch with
sterigmata and oblong spores.
4. The _Squamaria_ type (called by Glück _Placodium_): also a branching
sporophore but with long sickle-like bent spores.
5. The _Parmelia_ type: a more complicated system of branching and
anastomosing of the sporophores, with oblong spores.
6, and 7. The _Sticta_ and _Physcia_ types: in both of these the
sporophores are multi-septate; they consist of a series of radiately
arranged hyphae rising from a basal tissue all round the pycnidium. They
anastomose to form a network and bud off “spermatia” from the free cells
or rather from minute sterigmata. In the _Physcia_ type there is more
general anastomosis of the sporophores and frequently masses of sterile
cells along with the fertile members occupy the centre of the pycnidium.
The spermatia of these and the following _Endocarpon_ type are short
cylindrical bodies (Fig. 113 B).
[Illustration: Fig. 113 B. 7, _Physcia ciliaris_ DC. × 600; 8,
_Endocarpon_ sp. × 600 (after Glück).]
8. _Endocarpon_ type: the pycnidium is filled by a tissue of short broad
cells, with irregular hollow spaces lined by fertile cells similar to
those of the _Sticta_ and _Physcia_ types.
The three last named types of sporophores represent Nylander’s section
of arthrosterigmata. Steiner has followed Nylander in also arranging the
various forms into two leading groups. The first, characterized by the
secondary branch or “sterigma,” he designates “exobasidial”; the second,
comprising the three last types in which the spores are borne directly on
the cells of the sporophore or on very short processes, he describes as
“endobasidial.” Steiner also introduces a new term, _fulcrum_, for the
sporophore.
The pycnidia in which these different sporophores occur are not, as a
rule, characteristic of one family. _Peltigera_ type is found only in
one family and the Cladonia type is fairly constant in _Cladoniae_,
but “_Psora_” pycnidia are found on very varying lichens among the
Lecideaceae, Verrucariaceae and others. The _Squamaria_ type with long
bent spores is found not only in _Squamaria_ (Glück’s _Placodium_) but
also in _Lecidea_, _Roccella_, _Pyrenula_, etc. _Parmelia_ type is
characteristic of many _Parmeliae_ and also of species of _Evernia_,
_Alectoria_, _Platysma_ and _Cetraria_. The _Sticta_ type occurs in
_Gyrophora_, _Umbilicaria_, _Nephromium_ and _Lecanora_ as well as in
_Sticta_ and in one species at least of _Collema_. To the _Physcia_ type
belong the pycnidia of most _Physciaceae_ and of various _Parmeliae_, and
to the closely related _Endocarpon_ type the pycnidia of _Endocarpon_ and
of _Xanthoria parietina_.
[Illustration: Fig. 114. Sterile filaments in spermogonia of _Lecidea
fuscoatra_ Ach. much magnified (after Lindsay).]
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