Even the larger lichens are attacked by these quick-growing crusts.
_Pertusaria globulifera_ spreads over _Parmelia perlata_ and _P.
physodes_, gradually dissolving and consuming the different thalline
layers; the lower cortex of the victim holds out longest and can be seen
as an undigested black substance within the _Pertusaria_ thallus for some
time. As a rule, however, the lichens with large lobes grow over the
smaller thalli in a purely mechanical fashion.
_c._ FROM OTHER VEGETATION. Zukal[861] has given instances of association
between mosses and lichens in which the latter seemed to play the part of
parasite. The terricolous species _Baeomyces rufus_ (_Sphyridium_) and
_Biatora decolorans_, as well as forms of _Lepraria_ and _Variolaria_,
he found growing over mosses and killing them. Stems and leaves of the
moss _Plagiothecium sylvaticum_ were grown through and through by the
hyphae of a _Pertusaria_, and he observed a leaf of _Polytrichum commune_
pierced by the rhizinae of a minute _Cladonia_ squamule. The cells had
been invaded and the neighbouring tissue was brown and dead.
Perhaps the most voracious consumer of organic remains is _Lecanora
tartarea_, more especially the northern form _frigida_. It is the
well-known cudbear lichen of West Scotland, and is normally a rock
species. It has an extremely vigorous thickly crustaceous and
quick-growing thallus, and spreads over everything that lies in its
path—decaying mosses, dead leaves, other lichens, etc. Kihlman[862] has
furnished a graphic description of the way it covers up the vegetation
on the high altitudes of Russian Lapland. More than any other plant it
is able to withstand the effect of the cold winds that sweep across
these inhospitable plains. Other plant groups at certain seasons or in
certain stages of growth are weakened or killed by the extreme cold of
the wind, and, immediately, a growth of the more hardy grey crust of
_Lecanora tartarea_ begins to spread over and take possession of the area
affected—very frequently a bank of mosses, of which the tips have been
destroyed, is thus covered up. In the same way the moorland _Cladoniae_,
_C. rangiferina_ (the reindeer moss) and some allied species, are
attacked. They have no continuous cortex, the outer covering of the long
branching podetia being a loose felt of hyphae; they are thus sensitive
to cold and liable to be destroyed by a high wind, and their stems,
which are blackened as decay advances, become very soon dotted with the
whitish-grey crust of the more vigorous and resistant _Lecanora_.
III. ASSIMILATION AND RESPIRATION
A. INFLUENCE OF TEMPERATURE
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