In _Coenogonium_, the alga is some species of _Trentepohlia_, a
filamentous genus mostly yellow, which often gives its colour to the
slender lichen filaments, the covering hyphae being very scanty. Other
filamentous species, such as _Usnea barbata_, etc., are persistently
greenish from the bright-green Protococcaceous cells lying near the
surface of the thalline strands. Many of the furfuraceous lichens are
greenish from the same cause, especially when moist, as are also the
larger lichens, _Physcia ciliaris_, Stereocaulons, Cladonias and others.
_b._ COLOUR DUE TO LICHEN-ACIDS. These substances, so characteristic of
lichens, are excreted from the hyphae, and lie in crystals on the outer
walls; they are generally most plentiful on exposed tissues such as
the cortex of the upper surface or the discs of the apothecia. Many of
these crystals are colourless and are without visible effect, except in
sometimes whitening the surface, strikingly exemplified in _Thamnolia
vermicularis_[884]; but others are very brightly coloured. These
latter belong to two chemical groups and are found in widely separated
lichens[885]:
1. Derivatives of pulvinic acid which are usually of a bright-yellow
colour. They are the colouring substance of _Letharia vulpina_, a
northern species, not found in our islands, of _Cetraria pinastri_ and
_C. juniperina_[886] which inhabit mountainous or hilly regions. The
crustaceous species, _Lecidea lucida_ and _Rhizocarpon geographicum_, owe
their colour to rhizocarpic acid.
The brilliant yellow of the crusts of some species of Caliciaceae is due
to the presence of the substance calycin, while coniocybic acid gives
the greenish sulphur-yellow hue to _Coniocybe furfuracea_. Epanorin
colours the hyphae and soredia of _Lecanora epanora_ a citrine-yellow and
stictaurin is the deep-yellow substance found in the medulla and under
surface of _Sticta aurata_ and _S. crocata_.
2. The second series of yellow acids are derivatives of anthracene. They
include parietin, formerly described as chrysophanic acid, which gives
the conspicuous colour to _Xanthoriae_ and to various wall lichens;
solorinic acid, the crystals of which cover the medullary hyphae and
give a reddish-grey tone to the upper cortex of _Solorina crocea_, and
nephromin which similarly colours the medulla of _Nephromium lusitanicum_
a deep yellow, the colour of the general thallus being, however, scarcely
affected. In this group must also be included the acids that cause
the yellow colouring of the medulla in _Parmelia subaurifera_ and the
yellowish thallus of some _Pertusariae_.
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