The presence of these coloured substances enables the cell-wall to resist
chemical reactions induced by the harmful influences of the atmosphere
or of the substratum. The darker the cell-wall and the more abundant
the pigment, the less easily is the plant injured either by acids or
alkalies. The coloured tips of the paraphyses thus give much needed
protection to the long lived sporiferous asci, and the dark thalline
tissues prevent premature rotting and decay.
_d._ ENUMERATION OF AMORPHOUS PIGMENTS:
=1. Green.= Bachmann found several different green pigments:
“Lecidea-green,” colouring red with nitric acid, is the dark blue-green
or olive-green (smaragdine) of the paraphyses of many apothecia in
the Lecideaceae, and may vary to a lighter blue; it appears almost
black in thalline cells[889]. “Aspicilia-green” occurs in the thalline
margin and sometimes in the epithecium of the fruits of species of
_Aspicilia_; it becomes a brighter green on the application of nitric
acid. “Bacidia-green,” also a rare pigment, becomes violet with the
same acid; it is found in the epithecium of _Bacidia muscorum_ and
_Bacidia acclinis_ (Lecideaceae). “Thalloidima-green” in the apothecia
of some species of _Biatorina_ is changed to a dirty-red by nitric acid
and to violet by potash. Still another termed “rhizoid-green” gives
the dark greenish colour to the rhizoids of _Physcia pulverulenta_
and _P. aipolia_ and to the spores of some species of _Physcia_ and
_Rhizocarpon_. It becomes more olive-green with potash.
=2. Blue.= A very rare colour in lichens, so far found in only a few
species, _Biatora_ (_Lecidea_) _atrofusca_, _Lecidea sanguinaria_ and
_Aspicilia flavida_ f. _coerulescens_. It forms a layer of amorphous
granules embedded in the outer wall of the paraphyses, becoming more
dense towards the epithecium. A few granules are also present in the
hymenium.
=3. Violet.= “Arthonia-violet” as it is called by Bachmann is a
constituent of the tissues of _Arthonia gregaria_, occurring in minute
masses always near the cortical cells; it is distinct from the bright
cinnabarine granules present in every part of the thallus.
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