Peirce[919] found that _Ramalina reticulata_ (Fig. 65), of which the
fronds are an open network, was mainly distributed by the tearing of the
lichen in high wind. This takes place during the winter rains, when not
only the lichen is wet and soft in texture, but when the deciduous trees
are bare of leaves, at a season, therefore, when the drifting thalline
scraps can again catch on to branch or stem. A series of observations on
the dispersal of forms of long pendulous Usneas was made by Schrenk[920].
In the Middle and North Atlantic States of America these filamentous
species rarely bear apothecia. The high winds break and disperse them
when they are in a wet condition. They generally grow on Spruces and
Firs, because the drifting filaments are more easily caught and entangled
on short needles. The successive wetting and drying causes them to coil
and uncoil, resulting in a tangle impossible to unravel, which holds them
securely anchored to the support.
D. ERRATIC LICHENS
In certain lichens, there is a tendency for the thallus to develop
excrescences of nodular form which easily become free and drift about
in the wind while still living and growing. They are carried sometimes
very long distances, and fall in thick deposits over localities far
from their place of origin. The most famous instance is the “manna
lichen,” _Lecanora esculenta_, which has been scientifically examined
and described by Elenkin[921]. He distinguishes seven different forms
of the species: f. _esculenta_, f. _affinis_, f. _alpina_, and f.
_fruticulosa-foliacea_ which are Alpine lichens, the remainder, f.
_desertoides_, f. _foliacea_ and f. _esculenta-tarquina_, grow on the
steppes or in the desert[922].
Elenkin[921] adds to the list of erratic lichens a variety of _Parmelia
molliuscula_ along with _P. ryssolea_ from S. Russia, from the Asiatic
steppes and from Alpine regions. Mereschkovsky[923] has also recorded
from the Crimea _Parmelia vagans_, probably derived from _P. conspersa_
f. _vaga_ (f. nov.). It drifts about in small rather flattened bits, and,
like other erratics, it never fruits.
Meyer[924] long ago described the development of wandering lichens:
scraps that were torn from the parent thallus continued to grow if there
were sufficient moisture, but at the same time undergoing considerable
change in appearance. The dark colour of the under surface disappears
in the frequently altered position, as the lobes grow out into narrow
intermingling fronds forming a more or less compact spherical mass;
the rhizoids also become modified and, if near the edge, grow out into
thread-like structures which bind the mass together. Meyer says that
“wanderers” have been noted as belonging to _Parmelia acetabulum_,
_Platysma glaucum_ and _Anaptychia ciliaris_.
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