_Urceolaria_ (_Diploschistes_) has a somewhat bulky crustaceous thallus
which may be almost evanescent in its semi-parasitic condition, the only
gonidia retained being in the margin of the apothecia. Nylander[939]
found isolated apothecia growing vigorously on _Cladonia_ squamules.
Hue[940] describes _Lecanora aspidophora_ f. _errabunda_, an Antarctic
lichen, as not only a wanderer but as a “shameless robber.” It is to be
seen everywhere on and about other lichens, settling small glomeruli of
apothecia here and there on the thallus of _Umbilicariae_ or between
the areolae of _Buelliae_, and always too vigorous to be ousted from its
position.
_Bacidia flavovirescens_ has been regarded by some lichenologists[941] as
a parasite on _Baeomyces_, but recent work by Tobler[942] seems to have
proved that the bright green thallus is that of the _Bacidia_.
_c._ PARASYMBIOSIS. There are certain lichens that are obligative
parasites and pass their whole existence on an alien thallus. They may
possibly have degenerated from the condition of facultative parasitism
as the universal history of parasitism is one of increased dependence
on the host, and of growing atrophy of the parasite, but, in the case
of lichens, there is always the peculiar symbiotic condition to be
considered: the parasite produces its own vigorous hyphae and normal
healthy fruits, it often claims only a share of the carbohydrates
manufactured by the gonidia. The host lichen is not destroyed by this
parasymbiosis though the tissues are very often excited to abnormal
growth by the presence of the invading organism.
Lauder Lindsay[943] was one of the first to study these “microlichens”
as he called them, and he published descriptions of those he had himself
observed on various hosts. He failed however to discriminate between
lichens and parasitic fungi. It is only by careful research in each
case that the affinity to fungi or to lichens can be determined; very
frequently the whole of them, as possessing no visible thallus, have been
classified with fungi, but that view ignores the symbiosis that exists
between the hyphae of the parasite and the gonidia of the host.
Parasitic lichens are rather rare on gelatinous thalli; but even among
these, a few instances have been recorded. Winter[944] has described a
species of _Leptoraphis_, the perithecia of which are immersed in the
thallus of _Physma franconicum_. The host is wholly unaffected by the
presence of the parasite except for a swelling where it is situated.
The foreign hyphae are easily distinguishable; they wander through the
thallus of the host with their free ends in the mucilage of the gonidial
groups from which they evidently extract nourishment. Species of the
lichen genus _Obryzum_ are also parasitic on gelatinous lichens.
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