In the following necessarily brief account of families and genera, the
“spermogonia” or “pycnidia” have in most cases been left out of account,
as in many instances they vary within the family and occasionally even
within the genus. Their taxonomic value is not without importance,
but, in the general systematic arrangement, they are only subsidiary
characters. An account of them has already been given, and for more
detailed statements the student is referred to purely systematic works.
There are two main types of spore production in the “pycnidia” which
have been shortly described by Steiner[1034] as “exobasidial” and
“endobasidial.” In the former the sporophores are simple or branched
filaments, at the apices of which a short process grows out and buds off
a pycnidiospore; in the latter the spores are budded directly from cells
lining the walls or filling the cavity of the pycnidium. The exobasidial
type is more simply rendered in the following pages by “acrogenous,”
the endobasidial by “pleurogenous” spore production. In many cases the
“spermogonia” or “pycnidia” are still imperfectly known. In designating
the gonidial algae, the more comprehensive Protococcaceae has been
substituted for _Protococcus_, as in many cases the alga is probably not
_Protococcus_ as now understood, but some other genus of the family[1035].
SUBCLASS I. ASCOLICHENS
SERIES I. PYRENOCARPINEAE
It is on mycological grounds that Pyrenocarpineae are placed at the base
of lichen classification. There is no evidence that the series was first
in time.
I. _MORIOLACEAE_
This family was described by Norman[1036] in 1872 from specimens
collected by himself in Norway or in the Tyrol, on soil or more
frequently on trees. There seems to have been no further record, and
Zahlbruckner, while accepting the family, suggests that an examination or
revision may be necessary.
The thallus is crustaceous. The algal cells, Protococcaceae, occur either
in groups (sometimes stalked) surrounded by a plectenchymatous wall
and called by Norman “goniocysts,” or they form nests in the thallus
termed “nuclei” which are surrounded by a double wall of plectenchyma,
colourless in the interior and brown outside. Norman invented the term
“Allelositismus,” which may be rendered “mutualism,” to indicate this
peculiar form of thallus. The species of _Spheconisca_ are fairly
numerous on poplars, willows and conifers:
Algae in “goniocysts” 1. *Moriola Norm.[1037]
Algae in double-walled “nuclei” 2. *Spheconisca Norm.
II. _EPIGLOEACEAE_
The family consists of but one genus and one species, _Epigloea
bactrospora_, and, according to Zahlbruckner, further examination is
necessary to make certain as to the lichenoid nature of the plant.
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