Cortex fastigate, of palisade hyphae.
Spores colourless.
Hypothecium black-carbonaceous.
Apothecia round.
Thallus fruticose 1. Roccella DC.
Thallus crustaceous-fruticose 2. *Roccellina Darbish.
Apothecia lirellate 3. *Reinkella Darbish.
Hypothecium colourless.
Gonidia present under the
hypothecium 4. *Pentagenella Darbish.
Gonidia absent from hypothecium 5. *Combea De Not.
Spores brown or brownish.
Medulla of parallel somewhat loose
hyphae 6. *Schizopelte Th. Fr.
Medulla solid, black 7. *Simonyella Steiner.
Cortex fibrous, of parallel hyphae.
Apothecia round.
Hypothecium black-carbonaceous.
Apothecia with thalline margin 8. *Dendrographa Darbish.
Apothecia with proper margin 9. *Roccellaria Darbish.
Hypothecium colourless 10. *Darbishirella A. Zahlbr.
Apothecia lirellate 11. *Ingaderia Darbish.
SUBSERIES 3. =Cyclocarpineae=
This last subseries includes the remaining twenty-nine families of
Ascolichens. They are very varied both in the fungal and the algal
symbionts. The fruit is more or less a discoid open apothecium. The
gonidia belong to different genera of Myxophyceae and Chlorophyceae,
but the most frequent are Protococcaceae. Families are based largely on
thalline structure.
XXIII. _LECANACTIDACEAE_
By many systematists this family is included under Graphidineae
on account of the fruit structure which in some of the forms is
carbonaceous and almost lirellate, and also because the algal symbiont is
_Trentepohlia_. The thallus is primitive, being thinly crustaceous and
non-corticate; the apothecium has a black carbonaceous hypothecium in two
of the genera, _Lecanactis_ and _Schismatomma_ (_Platygrapha_); in the
third genus, _Melampydium_, it is colourless. The latter is monotypic,
and the spores become muriform. In the other genera they are elongate and
multi-septate.
Apothecia with prominent proper margin 1. Lecanactis Eschw.
Apothecia with thin proper margin 2. *Melampydium Müll.-Arg.
Apothecia with thalline margin 3. Schismatomma Flot.
XXIV. _PILOCARPACEAE_
A small family with but one genus, _Pilocarpon_. It is distinguished
as one of the few epiphyllous genera of lichens associated with
Protococcaceous gonidia and with a distribution extending far beyond
the tropics. The best known species, _P. leucoblepharum_, encircles the
base of pine-needles with a white felted crust, or inhabits coriaceous
evergreen leaves. Another species lives on fern leaves. The fruit is
a discoid apothecium with a dark carbonaceous hypothecium and proper
margin, and with a second thalline margin. The paraphyses are branched
and interwoven above.
Spores elongate, 3-septate, colourless 1. Pilocarpon Wain.
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